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A0163 Hesperocyparis arizonica Forest & Woodland Alliance

Type Concept Sentence: This forest and woodland alliance is dominated by Hesperocyparis arizonica with Juniperus deppeana, Pinus discolor, Quercus arizonica, Quercus palmeri, or Quercus hypoleucoides present to codominant in the subcanopy. It is found in desert mountain ranges in southeastern Arizona, southern New Mexico, and northern Mexico.


Common (Translated Scientific) Name: Arizona Cypress Forest & Woodland Alliance

Colloquial Name: Arizona Cypress Forest & Woodland

Hierarchy Level:  Alliance

Type Concept: This forest and woodland alliance is found in scattered locations in "Sky Island" mountain ranges in southeastern Arizona and southern New Mexico, and in the Sierra Madre Occidentale in Chihuahua and Sonora, Mexico. The open to closed tree canopy is dominated by Hesperocyparis arizonica. Other trees are often present to codominant include Juniperus deppeana, Pinus discolor, Quercus arizonica, Quercus palmeri, or Quercus hypoleucoides. The shrub layer is heavy and diverse. Common shrubs and rosette stem succulents include Arctostaphylos pungens, Ceanothus fendleri, Cercocarpus montanus, Fendlera rupicola, Fraxinus anomala, Garrya wrightii, Nolina microcarpa, Prunus virginiana, Quercus rugosa, and Quercus turbinella. The herbaceous layer is also diverse with many graminoids such as Bouteloua curtipendula, Carex geophila, Elymus elymoides, Koeleria macrantha, Muhlenbergia longiligula, Muhlenbergia richardsonis, Piptochaetium pringlei, Poa fendleriana, and Schizachyrium scoparium. The forb Packera neomexicana is common. Fires may be important for Hesperocyparis arizonica seed to germinate. This forest and woodland alliance is found on slopes and in drainages. Sites tend to be relatively cool and moist mountain slopes and canyons between 1220 and 2000 m elevation with north or east aspects. Substrates are shallow, rocky soil.

Diagnostic Characteristics: This forest and woodland alliance is characterized by an evergreen tree canopy dominated or codominated by Hesperocyparis arizonica trees. Associated tree species include Pinus discolor, Quercus hypoleucoides, Quercus palmeri, and Quercus arizonica.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Woodland stands dominated by Hesperocyparis nevadensis and Hesperocyparis stephensonii occur in southern California and Baja California, Mexico (Griffin and Critchfield 1976, Barbour and Major 1977). Both woodland types were classified and described as associations in Reid et al. (1994). These woodlands are currently not included in this alliance or any other. Hesperocyparis arizonica forest stands occurring in riparian zones along perennial streams need further review.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Vegetation included in this alliance has a moderately dense to dense evergreen tree canopy (35-100% cover), 5-20 m tall, dominated by needle-leaved evergreen trees. Often there are tall- and short-shrub layers dominated by broad-leaved evergreen species with broad-leaved deciduous and needle-leaved species present ranging from 0.5-5 m tall. The herbaceous layer is typically sparse and dominated by perennial graminoids, with annual forbs and grasses present seasonally.

Floristics: This alliance has a moderately dense to dense (>50% cover) tree canopy dominated by the evergreen conifer Hesperocyparis arizonica (= Cupressus arizonica). The broad-leaved tree Quercus hypoleucoides codominates some stands. Other trees are often present to codominant, including conifers Juniperus deppeana, Juniperus monosperma, Pinus edulis, and Pinus monophylla (= Pinus fallax), and evergreen oaks such as Quercus arizonica, Quercus palmeri, and Quercus oblongifolia. The shrub layer is dense and diverse often with over 12 species present. Quercus turbinella, a deciduous broad-leaved shrub, is the most important and a diagnostic species. Other common shrubs and rosette stem succulents include Arctostaphylos pungens, Ceanothus fendleri, Cercocarpus montanus, Fendlera rupicola, Fraxinus anomala, Garrya wrightii, Nolina microcarpa, Prunus virginiana, and Quercus rugosa. The herbaceous layer is typically sparse (<5% cover) because of shading by the overstory, but may be moderately dense depending on the density of the woody canopies and moisture. It is dominated by diverse graminoids such as Bouteloua curtipendula, Carex geophila, Elymus elymoides (= Sitanion hystrix), Koeleria macrantha, Muhlenbergia longiligula, Muhlenbergia richardsonis, Piptochaetium pringlei (= Stipa pringlei), Poa fendleriana, and Schizachyrium scoparium. Packera neomexicana (= Senecio neomexicanus) and Galium spp. are typical of the sparse forb layer. Carmichael et al. (1978) described stands in the Mazatzal Mountains with average cover of Hesperocyparis arizonica, Quercus turbinella, and 11 other shrubs being 37%, 26% and 10%, respectively, for total average cover of 73%.

Dynamics:  Stands occur on a variety of substrates which indicates that other factors, most likely climatic conditions, restrict the distribution of this alliance. The cool, moist conditions required by Hesperocyparis arizonica are not common in southern Arizona. Few occurrences of this alliance have been documented. Brown (1982a) considers these stands to be relictual. Parker (1980b) thinks Hesperocyparis arizonica is a stable, terminal element of limited environments where it now occurs. Its life history resembles that of a pioneer species with additional characteristics that allow it to persist. A trait that leads to persistence is that Hesperocyparis arizonica is very long-lived. The extrapolated ages of individuals from one stand averaged 317 years with the oldest tree being 456 years (Parker 1980b). Also, known stands tend to be mature with little regeneration, and seedlings are able to germinate and grow in the shade of mature trees. Pioneer traits are that it is a prolific seed-producer (estimated 0.1-1 million seeds per tree) and intolerant of ground litter (Parker 1980b). The two stands with a significant number of seedlings had both been disturbed, one by logging and one by flood. Moist, mineral soil is the ideal seedbed for Hesperocyparis arizonica. Parker (1980b) found the number of seedlings decreases with increased litter layer depth.

Fire appears to be important for Hesperocyparis arizonica stand regeneration. Carmichael et al. (1978) report fire opens Hesperocyparis arizonica seed cones. Although Hesperocyparis arizonica is found in areas that are protected from fire and is a non-sprouter with limited fire adaptation, fire removes accumulations of ground litter that prevent seedling establishment (Parker 1980a, 1980b). Parker (1980b) suggested that fire suppression may have limited reproduction, especially in habitats that do not flood, and that current fire suppression policies be eliminated in areas where Hesperocyparis arizonica occurs.

Environmental Description:  This alliance includes relict forests that are restricted to local mesic sites in mountains in southeastern Arizona, and a site in southwestern New Mexico. Elevation ranges from 1220-2000 m depending on aspect. Climate is arid to semi-arid with temperatures rarely falling below freezing. Annual precipitation has a bimodal distribution with about half of the 50 cm mean annual precipitation occurring in July through September during the late summer monsoon, and most of the rest falling during the winter months. Where sites have less precipitation or warmer aspects, soil moisture is usually augmented by a spring or stream. Stands of Hesperocyparis arizonica forest occur primarily on north- and east-facing mountain slopes and canyons, and along drainages where both summer and winter temperatures are moderate. Sites range from gentle alluvial toeslopes to steep canyon and mountain slopes (3-60%). Stands are generally small and found in "cool" pockets within warm Madrean evergreen woodland or chaparral where local edaphic conditions permit a shallow water table. Substrate is derived from a variety of parent materials, but soils are generally rocky or gravelly loams or sandy loams. Hesperocyparis arizonica has also been reported growing on drier sites at elevations above 2050 m, but stands are sparse and stunted, and resemble a sparse shrub savanna (Parker 1980a).

Geographic Range: Forests included in this alliance occur locally on mesic slopes and drainages in sub-Mogollon Arizona in the Blue Range, Dragoon, Santa Catalina, Mazatzal and Chiricahua mountains, and in one location in southwestern New Mexico near Cooke''s Peak in the Black Range. Its range extends into northern Mexico along the Sierra Madre Occidentale in Chihuahua and Sonora. The alliance may also occur in the Big Bend region of Texas and Mexico (Parker 1980a, 1980b).

Nations: MX,US

States/Provinces:  AZ, MXCHH, MXSON, NM, TX?




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: Old Alliance I.A.8.N.c. Cupressus arizonica Forest Alliance (A.163) is old name.

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Cupressus arizonica/Quercus hypoleucoides Habitat Type (Bassett et al. 1987)
? Cupressus arizonica/Quercus turbinella Habitat Type (Bassett et al. 1987)
? Cupressus arizonica association (Brown et al. 1979) [within the Relict Conifer Forest and Woodland Cypress Series (123.52) Cupressus arizonica association]
? Cypress Canyon (Niering and Lowe 1984)

Concept Author(s): D.E. Brown, C.H. Lowe and C.P. Pase (1979)

Author of Description: K.A. Schulz and M.S. Reid

Acknowledgements: No Data Available

Version Date: 01-08-14

  • Barbour, M. G., and J. Major, editors. 1977. Terrestrial vegetation of California. John Wiley and Sons, New York. 1002 pp.
  • Barrows, J. S., E. W. Mogren, K. Rowdabaugh, and R. Yancik. 1977. The role of fire in ponderosa pine and mixed conifer ecosystems. Final report, Cooperative report between the National Park Service and Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 101 pp.
  • Bassett, D., M. Larson, and W. Moir. 1987. Forest and woodland habitat types of Arizona south of the Mogollon Rim and southwestern New Mexico. Edition 2. USDA Forest Service, Southwestern Region, Albuquerque, NM.
  • Brown, D. E., C. H. Lowe, and C. P. Pase. 1979. A digitized classification system for the biotic communities of North America with community (series) and association examples for the Southwest. Journal of the Arizona-Nevada Academy of Science 14:1-16.
  • Brown, D. E., editor. 1982a. Biotic communities of the American Southwest-United States and Mexico. Desert Plants Special Issue 4(1-4):1-342.
  • Carmichael, R. S., O. D. Knipe, C. P. Pase, and W. W. Brady. 1978. Arizona chaparral: Plant associations and ecology. Research Paper RM-202. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 16 pp.
  • Faber-Langendoen, D., J. Drake, M. Hall, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, M. Russo, K. Schulz, L. Sneddon, K. Snow, and J. Teague. 2013-2019b. Screening alliances for induction into the U.S. National Vegetation Classification: Part 1 - Alliance concept review. NatureServe, Arlington, VA.
  • Griffin, J. R., and W. B. Critchfield. 1976. The distribution of forest trees in California. Research Paper PSW-82. USDA Forest Service, Pacific Southwest Forest and Range Experiment Station, Berkeley, CA. 115 pp.
  • Larson, M., and W. H. Moir. 1986. Forest and woodland habitat types (plant associations) of southern New Mexico and central Arizona (north of the Mogollon Rim). USDA Forest Service, Southwestern Region, Albuquerque, NM. 76 pp.
  • Niering, W. A., and C. H. Lowe. 1984. Vegetation of the Santa Catalina Mountains: Community types and dynamics. Vegetatio 58:3-28.
  • Parker, A. J. 1980a. The successional status of Cupressus arizonica. Great Basin Naturalist 40(3):254-264.
  • Parker, A. J. 1980b. Site preferences and community characteristics of Cupressus arizonica Greene (Cupressacaeae) in southeastern Arizona. Southwestern Naturalist 25(1):9-22.
  • Reid, M. S., L. S. Engelking, and P. S. Bourgeron. 1994. Rare plant communities of the conterminous United States, Western Region. Pages 305-620 in: D. H. Grossman, K. L. Goodin, and C. L. Reuss, editors. Rare plant communities of the conterminous United States, an initial survey. The Nature Conservancy, Arlington, VA.