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A0537 Pinus aristata Woodland Alliance
Type Concept Sentence: This woodland alliance with canopies dominated by Pinus aristata occurs on semi-xeric exposed sites in the subalpine zone throughout the southern Rocky Mountains and eastern Colorado Plateau.
Common (Translated Scientific) Name: Bristlecone Pine Woodland Alliance
Colloquial Name: Bristlecone Pine Woodland
Hierarchy Level: Alliance
Type Concept: This woodland alliance occurs on semi-xeric sites in the subalpine zone throughout the southern Rocky Mountains and eastern Colorado Plateau. Woodlands included in this alliance are characterized by an open to moderately closed tree canopy typically 5-15 m tall that is dominated by the conifer Pinus aristata. Under the harshest conditions it may occur as krummholz composed of stunted individuals. In some stands trees are clumped with grassy patches interspersed. Other tree species that may be present to codominant include Picea engelmannii, Pseudotsuga menziesii, and less commonly Abies lasiocarpa, Picea pungens, Pinus flexilis, and Populus tremuloides. The understory vegetation ranges from moderately dense to typically sparse because sites are dry and often have large amounts of rock cover. The shrub layer is generally sparse or absent, and can include Dasiphora fruticosa ssp. floribunda, Juniperus communis, Ribes spp., Rosa woodsii, Saxifraga bronchialis, Symphoricarpos oreophilus, and immature trees. The sparse to moderately dense herbaceous layer often dominates the understory. The most common species are typically graminoids, especially species of Festuca. Other common species may include Calamagrostis purpurascens, Carex spp., Danthonia parryi, Festuca arizonica, Festuca brachyphylla, Festuca idahoensis, Festuca thurberi, Koeleria macrantha, Muhlenbergia filiculmis, Muhlenbergia montana, Poa fendleriana, and Trisetum spicatum. Forbs are generally sparse. Sites are typically xeric on exposed, windswept rocky slopes and ridges. Stands occur on all aspects, but are most common on the drier south- and west-facing slopes. Slopes are variable. Soils are typically well-drained, shallow, skeletal and coarse-textured.
Diagnostic Characteristics: Woodlands of the subalpine zone throughout the southern Rocky Mountains and eastern Colorado Plateau dominated or codominated by Pinus aristata. Typically these sites are extremely harsh and only Pinus aristata is competitive enough to occupy these environments, therefore codominant species are rare.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: Some Pinus aristata stands may have Pinus flexilis as an associate (Rominger and Paulik 1983, Larson and Moir 1987, Komarkova et al. 1988a, Ranne et al. 1997). However, Pinus aristata is rarely found in stands dominated by Pinus flexilis because Pinus aristata appears to be more competitive and dominates these sites (Peet 1978, 1981). These woodlands are typically open, but stands on scree slopes, especially those in ~Pinus aristata / Ribes montigenum Woodland (CEGL000761)$$, have too sparse a tree canopy to be classified as a woodland (Shepherd 1975, Johnson 1987, Larson and Moir 1987). Ranne et al. (1997) described two additional associations that need to be reviewed for possible inclusion into the National Vegetation Classification.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: Vegetation included in this alliance has an open tree canopy that is typically up to 5-15 m tall. The canopy is dominated by evergreen needle-leaved trees. Sparse to moderately sparse shrub layers may be present. If present, the shrub layer is typically dominated by broad-leaved deciduous or evergreen scale-leaved shrubs usually less than 2 m tall. A sparse to moderately dense herbaceous layer dominated by perennial graminoids is often present. Perennial forbs have sparse cover. Annual forbs and grasses may be seasonally present.
Floristics: Stands have an open to moderately closed canopy typically 5-15 m tall that is solely dominated or codominated by the long-lived evergreen needle-leaved tree Pinus aristata. Individual trees may reach 20 m. In some stands trees are clumped with grassy patches interspersed. Other tree species that may be present to codominant vary geographically within its range. Picea engelmannii and Pseudotsuga menziesii are the most common. Picea pungens is common in the southern extent where stands occur at high elevations, and Pinus flexilis has also been reported as an associate in some stands. Abies lasiocarpa and Populus tremuloides may be scattered in some stands, but are generally restricted to more mesic sites and are typically absent in the drier southern extent of this alliance. The understory vegetation ranges from moderately dense to typically sparse because sites are dry and often have large amounts of rock cover. The shrub layer is generally sparse or absent. If present, shrubs are generally scattered, except for Juniperus communis, which often occurs in clumps. Other shrub species may include Dasiphora fruticosa ssp. floribunda (= Pentaphylloides floribunda), Ribes cereum, Ribes montigenum, Rosa woodsii, Saxifraga bronchialis, Symphoricarpos oreophilus, and immature trees. The sparse to moderately dense herbaceous layer often dominates the understory. The most common species are typically graminoids, especially species of Festuca. Other common species may include Calamagrostis purpurascens, Carex spp., Danthonia parryi, Festuca arizonica, Festuca brachyphylla, Festuca idahoensis, Festuca thurberi, Koeleria macrantha, Muhlenbergia filiculmis, Muhlenbergia montana, Poa fendleriana, and Trisetum spicatum. Forbs are generally sparse, but Trifolium dasyphyllum may reach 10% cover. Other scattered forbs may include species of Achillea, Antennaria, Artemisia, Arenaria, Arnica, Astragalus, Campanula, Erigeron, Hymenoxys, Penstemon, Polemonium, Sedum, Senecio, and Thalictrum.
Dynamics: Pinus aristata is a slow-growing, extremely long-lived tree (Brunstein and Yamaguchi 1992). Several individuals over 2000 years old have been found in Colorado. DeVelice et al. (1986) observed it occurs in open park-like stands and on steep rocky slopes where Picea engelmannii - Abies lasiocarpa stands are excluded by drought. Fire is important in the grass-dominated stands but is rarely intense enough to result in tree-killing crown fires (DeVelice et al. 1986). In some stands, suppression of grass fires has allowed encroachment of the trees into meadows (Larson and Moir 1987). Forage production is good in some stands, but rarely utilized by livestock because stands are steep and generally remote which makes access difficult. Peet (1978, 1981) noted that Pinus aristata is dominant at higher elevations in much of the southern Rocky Mountains, where Pinus flexilis is restricted to lower elevations. This is attributed to apparent competitive exclusion, because Pinus flexilis is dominant at high elevations in northern Colorado, Wyoming and Montana.
Environmental Description: Woodlands included in this alliance occur on semi-xeric sites in the subalpine zone throughout the southern Rocky Mountains and eastern Colorado Plateau. Elevations range from 2600-3670 m. Climate is semiarid, cold temperate with cool summers. Annual precipitation patterns and amounts vary with latitude, but locally the sites are typically xeric on exposed, windswept rocky slopes and ridges. These open to moderately closed woodlands occur on all aspects, but are most common on the drier south- and west-facing slopes. Slopes are typically moderate to steep, but may also be gentle. Soils are typically well-drained, shallow, skeletal and coarse-textured such as gravelly, sandy loams or loams. Stands occur most frequently on igneous, metamorphic and volcanic substrates such as andesite, granite, gneiss, breccia, tuff, conglomerate, but also occur on sedimentary rocks like sandstone. Exposed bedrock is common. Soil pH is 4.5-6.9, acidic to slightly acidic.
Geographic Range: Stands included in this woodland alliance occur intermittently throughout the southern Rocky Mountains and on mountains and plateaus on the eastern Colorado Plateau.
Nations: US
States/Provinces: AZ, CO, NM, UT?
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.898999
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Concept Lineage: A.537
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: ? Pinus aristata Series (Johnston 1987)
? Bristlecone Pine Series (Dick-Peddie 1993)
? Bristlecone Pine: 209 (Eyre 1980)
? Bristlecone Pine Series (Dick-Peddie 1993)
? Bristlecone Pine: 209 (Eyre 1980)
- Brunstein, C. R., and D. K. Yamaguchi. 1992. The oldest known Rocky Mountain bristlecone pines (Pinus aristata Engelm.). Arctic and Alpine Research 24:253-256.
- DeVelice, R. L., J. A. Ludwig, W. H. Moir, and F. Ronco, Jr. 1986. A classification of forest habitat types of northern New Mexico and southern Colorado. General Technical Report RM-131. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 59 pp.
- Dick-Peddie, W. A. 1993. New Mexico vegetation: Past, present, and future. University of New Mexico Press, Albuquerque. 244 pp.
- Eyre, F. H., editor. 1980. Forest cover types of the United States and Canada. Society of American Foresters, Washington, DC. 148 pp.
- Faber-Langendoen, D., J. Drake, M. Hall, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, M. Russo, K. Schulz, L. Sneddon, K. Snow, and J. Teague. 2013-2019b. Screening alliances for induction into the U.S. National Vegetation Classification: Part 1 - Alliance concept review. NatureServe, Arlington, VA.
- Johnson, K. L. 1987a. Sagebrush types as ecological indicators to integrated pest management (IPM) in the sagebrush ecosystem of North America. Pages 1-10 in: J. A. Onsager, editor. Integrated pest management on rangelands: State of the art in the sagebrush ecosystem. USDA Agricultural Research Service ARS-50. 85 pp.
- Johnston, B. C. 1987. Plant associations of Region Two: Potential plant communities of Wyoming, South Dakota, Nebraska, Colorado, and Kansas. R2-ECOL-87-2. USDA Forest Service, Rocky Mountain Region. Lakewood, CO. 429 pp.
- Komarkova, V., A. Peters, G. Kamani, W. Jones, V. Howard, H. Gordon, and K. Southwick. 1988a. Natural recovery of plant communities on disturbance plots and history of land use in the Niwot Ridge/Green Lakes Valley, Front Range, Colorado. University of Colorado Longterm Ecological Research Working Paper 88/1. Boulder, CO. 46 pp.
- Larson, M., and W. H. Moir. 1987. Forest and woodland habitat types of northern New Mexico and northern Arizona. Edition 2. USDA Forest Service, Southwestern Region, Albuquerque, NM.
- Peet, R. K. 1978. Latitudinal variation in southern Rocky Mountain forests. Journal of Biogeography 5:275-289.
- Peet, R. K. 1981. Forest vegetation of the Colorado Front Range. Vegetatio 45:3-75.
- Ranne, B. M., W. L. Baker, T. Andrews, and M. G. Ryan. 1997. Natural variability of vegetation, soils, and physiography in the bristlecone pine forests of the Rocky Mountains. Great Basin Naturalist 57(1):21-37.
- Rominger, J. M., and L. A. Paulik. 1983. A floristic inventory of the plant communities of the San Francisco Peaks Research Natural Area. General Technical Report RM-96. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 9 pp.
- Shepherd, H. R. 1975. Vegetation of two dissimilar bighorn sheep ranges in Colorado. Colorado Division of Wildlife Report 4. 223 pp.