Print Report

A0386 Quercus incana - Quercus arkansana - Pinus echinata Woodland Alliance

Type Concept Sentence: This alliance includes communities dominated by a combination of Pinus echinata (rarely with some Pinus taeda) with the scrubby oaks Quercus incana, Quercus margarettae, and/or Quercus arkansana. Quercus incana is the most characteristic and typically dominant of these. Stands of this alliance occur on xeric, sandy sites in the Coastal Plain west of the Mississippi River in eastern Texas, southeastern Oklahoma, northwestern Louisiana, and southwestern Arkansas.


Common (Translated Scientific) Name: Bluejack Oak - Arkansas Oak - Shortleaf Pine Woodland Alliance

Colloquial Name: West Gulf Coastal Plain Scrub Oak - Shortleaf Pine Sandhill Woodland

Hierarchy Level:  Alliance

Type Concept: This alliance includes communities dominated by a combination of Pinus echinata (rarely with some Pinus taeda) with the scrubby oaks Quercus incana, Quercus margarettae, and/or Quercus arkansana. Quercus incana is the most characteristic and typically dominant of these. In addition to the aforementioned species, Carya texana may be present. Some stands are short-statured, deciduous woodlands with scattered pines, while others are more pine-dominated and forest-like. A patchy shrub stratum (varying from approximately 30-60% in total cover) consisting of stunted, scrubby Quercus spp. and tall shrubs ranging from 2.5-5 m in height is usually present. Quercus arkansana, a rare Coastal Plain endemic species, may occur in some examples. Due to xeric conditions, graminoids and forbs are very sparse in this community, and patches of exposed sand are common. Lichens (Cladonia spp.) and Selaginella arenicola ssp. riddellii form large patches. Typical shrubs include Asimina parviflora, Chionanthus virginicus, Frangula caroliniana, Ilex vomitoria, Sassafras albidum, Sideroxylon lanuginosum ssp. lanuginosum, Stillingia sylvatica, Vaccinium arboreum, and seedlings of canopy species. Depending, in part, upon management history, the herbaceous layer density may vary widely and is sometimes quite sparse. Stands of this alliance occur on xeric, sandy sites in the Coastal Plain west of the Mississippi River in eastern Texas, southeastern Oklahoma, northwestern Louisiana, and southwestern Arkansas. The canopy coverage may be variable, reflecting the range of situations involved, ranging from open-canopied woodlands on very well-drained Pleistocene terraces and low, broad ridges on deep, acidic sandy soils to slightly more-or-less dense examples on similarly droughty sites.

Diagnostic Characteristics: These are dry to very dry (xeric) oak woodlands on sandy substrates in the West Gulf Coastal Plain of the southeastern United States.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This alliance encompasses some physiognomic variation, which is characteristic of this vegetation. This alliance is coherent with ~West Gulf Coastal Plain Sandhill Oak and Shortleaf Pine Forest and Woodland (CES203.056)$$. In regard to the floristic distinctiveness of G013 and its alliances (A0386, A3129, A3130), the available data do not reveal any specifically diagnostic or differential species in relation to similar alliances in groups from other parts of the southern coastal plains (Atlantic and East Gulf). There are, however, diagnostic and differential species that distinguish these alliances from one another.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Some stands are short-statured, deciduous woodlands, while others are more forest-like. Depending, in part, upon management history, the herbaceous layer density may vary widely and is sometimes quite sparse. The canopy coverage may be variable, reflecting the range of situations involved. Some associations may exhibit an open canopy of Pinus echinata (sometimes with Pinus taeda) ranging generally from 15-40% cover, while other examples are slightly more dense. A stunted, scrubby Quercus spp. and tall-shrub layer (2.5-5 m in height) is typically present. This stratum is patchy and varies from approximately 30-60% in cover. Due to xeric conditions, graminoids and forbs are very sparse in this community, and patches of exposed sand are common. Lichens (Cladonia spp.) and Selaginella arenicola ssp. riddellii may form large patches.

Floristics: Vegetation indicators are species tolerant of droughty sites, especially Quercus incana and Quercus arkansana (a rare Coastal Plain endemic species), but also Quercus marilandica and Quercus stellata. Pinus echinata is usually present in examples of this alliance, and Pinus palustris is typically absent (or perhaps at low frequency within its range). In addition to the aforementioned species, Carya texana may be present. In most cases, a patchy shrub stratum (varying from approximately 30-60% in total cover) consisting of stunted, scrubby Quercus spp. and tall shrubs ranging from 2.5-5 m in height is usually present. Among the subcanopy taxa, Quercus incana is arguably the most constant species. Typical shrubs include Asimina parviflora, Chionanthus virginicus, Frangula caroliniana, Ilex vomitoria, Sideroxylon lanuginosum ssp. lanuginosum, Stillingia sylvatica, Vaccinium arboreum, and seedlings of canopy species. Patches of exposed sand are common in this alliance. Due to xeric conditions and depending, in part, upon management history, the herbaceous layer density may vary widely and is sometimes quite sparse. However, in nearly all cases, it will contain a suite of habitat fidels, several of which are either endemic or nearly so to the West Gulf Coastal Plain. The widely scattered herbaceous layer may contain Aristida desmantha, Cnidoscolus texanus, Eriogonum longifolium, Eriogonum multiflorum, Opuntia humifusa, Penstemon murrayanus, Polanisia erosa, Polygonella americana, Polygonella polygama, Schizachyrium scoparium, and Zornia bracteata. More open examples that approach woodland structure may contain scattered patches of fruticose lichens (Cladonia spp.) and possibly Selaginella arenicola ssp. riddellii. The most extreme examples are open, xeric woodlands with a sparse canopy of the nominal species over open areas dominated by grass, bare sand, or Selaginella arenicola ssp. riddellii. Shrubs occur in dense patches and may include Carya texana, Chionanthus virginicus, Frangula caroliniana, Rhus aromatica, Sassafras albidum, Sideroxylon lanuginosum ssp. lanuginosum, Vaccinium arboreum, and Vaccinium stamineum. Herbaceous species may include Asclepias spp., Clematis sp., Delphinium sp., Oenothera heterophylla, Opuntia sp., Penstemon murrayanus, and Schizachyrium scoparium. This vegetation supports a large concentration of vascular plant endemics, near endemics, and a number of plant species with high fidelity to sandhills in the region.

Dynamics:  This alliance primarily includes natural woodlands and forests of the drier parts of the region, but some vegetation may be included here which has resulted from the effects of fire suppression. The historical fire frequency of the natural types is unknown but is widely believed to have been less than that of Pinus palustris-dominated woodlands (which sometimes occur adjacent to this type). Too frequent fires would not allow for the persistence of Pinus echinata and hardwood species, although young Pinus echinata has the ability to resprout when top-killed by fire. In some areas of eastern Texas, these xeric sandhills occupy very limited portions of the landscape, possibly suggesting that natural fire frequencies may not have been as constant or as frequent as would be necessary to maintain herbaceous-dominated woodlands. In addition, the extremely droughty soils contribute to only sparse fine fuel build-up making frequent, natural fires difficult to either ignite or spread. In the continued absence of fire, this mixed forest alliance may become increasingly invaded by Pinus taeda or may be seral to a xeric Quercus incana - Quercus margarettae - Quercus hemisphaerica-dominated forest.

Environmental Description:  Communities in this alliance occur on xeric uplands underlain with deep, coarse sandy soils, including very well-drained Pleistocene terraces and low, broad ridges on deep, acidic soils, as well as ridgetops and flat xeric uplands on very well-drained, acidic (often Grossarenic) sandy soils. These sites are typified by low fertility and moisture retention, which contribute to open tree canopies with usually <60% canopy closure. Sparse understory vegetation and abundant patches of bare soil are indicative of this alliance. Droughty conditions associated with these soils coupled with the effects of natural and managed fire contribute to extreme openness of the vegetation in some stands, which may appear to be an herbaceous-dominated community with scattered emergent woody stems in the most extreme cases.

Geographic Range: This alliance is restricted to the West Gulf Coastal Plain and Upper West Gulf Coastal Plain in eastern Texas, southeastern Oklahoma, northwestern Louisiana, and southwestern Arkansas.

Nations: US

States/Provinces:  AR, LA, OK, TX




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: (A.386)

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: No Data Available

Concept Author(s): M. Pyne, in Faber-Langendoen et al. (2013)

Author of Description: M. Pyne

Acknowledgements: We have incorporated significant descriptive information previously compiled by Rob Evans.

Version Date: 01-08-14

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