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G390 Psidium spp. - Syzygium jambos - Grevillea robusta Ruderal Lowland Rainforest Group

Type Concept Sentence: This Polynesian lowland ruderal group contains all mesic and wet forests and woodlands that are strongly dominated (>90% relative canopy cover) by non-native trees in the tree canopy such as Acacia confusa, Araucaria columnaris, Casuarina spp. (coastal areas), Coffea arabica, Grevillea robusta, Mangifera indica, Morella faya, Phyllostachys nigra, Phoenix dactylifera, Psidium cattleianum, Psidium guajava, Schefflera actinophylla, Schinus terebinthifolius, Syzygium cumini, Syzygium jambos, Thespesia populnea (common along coasts), and native tree canopy dominated or codominated by Acacia koa with exotic ruderal understory.


Common (Translated Scientific) Name: Guava species - Malabar Plum - Silk-oak Ruderal Lowland Rainforest Group

Colloquial Name: Polynesian Ruderal Lowland Rainforest

Hierarchy Level:  Group

Type Concept: This Polynesian lowland ruderal group contains all mesic and wet forests and woodlands that are strongly dominated (>90% relative canopy cover) by non-native trees in the tree canopy. Numerous species may dominate these disturbed stands, including Acacia confusa, Araucaria columnaris, Casuarina spp. (coastal areas), Coffea arabica, Eucalyptus spp., Grevillea robusta, Mangifera indica, Morella faya, Phyllostachys nigra, Phoenix dactylifera, Psidium cattleianum, Psidium guajava, Schefflera actinophylla, Schinus terebinthifolius, Syzygium cumini, Syzygium jambos, Thespesia populnea (common along coasts), and Trema orientale. Also included are stands with a tree canopy dominated or codominated native species, such as Acacia koa or Metrosideros polymorpha, with a severely degraded understory that is altered beyond the point that the natural plant community can be determined, and the community has converted to a ruderal or novel type (usually dominated by non-native species with>90% relative cover). There are numerous, invasive, exotic understory species, especially trees Psidium cattleianum and Morella faya, and herbaceous species Ageratum conyzoides, Andropogon virginicus, Axonopus fissifolius, Melinis minutiflora, Paspalum conjugatum, Sacciolepis indica, and Sphagneticola trilobata. Stands in this group occur in Hawai`i, Moloka`i, Maui, Kaua`i and O`ahu, and elsewhere in Polynesia. Sites occur along the coasts inland from 5 to 1500 m (15-4920 feet) elevation, but are generally found below 1000 m (3280 feet). Coastal forests also occur on windward sides where they intercept sufficient orographic rainfall to maintain a forest condition, yet may be influenced by coastal factors such as salt spray. The mesic forests have relatively even, moderate precipitation throughout the year or have a distinct, seasonally dry period and typically do not experience extended periods of drought like the dry forests and woodlands. Annual rainfall is 1200-2500 mm (47-98 inches) and falls largely from October to March. Rainforest are typically wet year round. Annual rainfall generally exceeds 2500 mm (98 inches). Substrates are highly variable but are generally well-drained and include steep, rocky talus, shallow to deep soils over weathered rock and gravelly alluvium, rocky shallow organic muck.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This ruderal group includes stands dominated by tree species that were introduced by Polynesians such as Aleurites moluccanus, Morinda citrifolia, Syzygium malaccense, and Thespesia populnea, recently (post-European contact) escaped ornamental and fruit trees Psidium spp., Melaleuca quinquenervia, and Morella faya, and non-native timber species that are invasive and naturalize in areas outside plantations. Although plantations may share some common timber species, the ruderal stands have often a multi-aged, mixed-species tree canopy rather than even-aged, near-monoculture of timber trees with clear stand plantation boundaries. Exotic timber species may include Acacia auriculiformis, Acacia confusa, Acacia mangium, Acacia mearnsii, Acacia melanoxylon, Albizia chinensis, Albizia lebbeck, Casuarina cunninghamiana, Eucalyptus globulus, Eucalyptus grandis, Eucalyptus saligna, Falcataria moluccana (= Albizia moluccana), Grevillea robusta, Pithecellobium dulce, and Toona ciliata and are generally treated as ruderal, unless found in plantations.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: This ruderal Polynesian group contains all mesic and wet forests and woodlands that are strongly dominated (>90% relative canopy cover) by non-native trees in the tree canopy such as Acacia confusa, Araucaria columnaris, Casuarina spp. (coastal areas), Coffea arabica, Eucalyptus spp., Grevillea banksii, Grevillea robusta, Mangifera indica, Morella faya, Phyllostachys nigra, Phoenix dactylifera, Psidium cattleianum, Psidium guajava, Schefflera actinophylla, Schinus terebinthifolius, Syzygium cumini, Syzygium jambos, Thespesia populnea (common along coasts), and Trema orientale. Also included are stands with a tree canopy dominated or codominated native species, such as Acacia koa or Metrosideros polymorpha, with a severely degraded understory that is altered beyond the point that the natural plant community can be determined, and the community has converted to a ruderal or novel type (usually dominated by non-native species with>90% relative cover). There are numerous, exotic understory species, including the especially invasive trees Psidium cattleianum and Morella faya, shrubs Caesalpinia decapetala, Lantana camara, and Triumfetta rhomboidea, and herbaceous species Ageratum conyzoides, Andropogon virginicus, Axonopus fissifolius, Cynodon dactylon, Melinis minutiflora, Paspalum conjugatum, Sacciolepis indica, and Sphagneticola trilobata.

Dynamics:  No Data Available

Environmental Description:  This lowland ruderal Polynesian forest and woodland group is found in mesic to wet areas of Hawai`i, Moloka`i, Maui, Kaua`i and O`ahu. Stands occur along the coasts inland from 5 to 1500 m (15-4920 feet) elevation, but are generally found below 1000 m (3280 feet). Coastal forests also occur on windward sides where they intercept sufficient orographic rainfall to maintain a forest condition, yet may be influenced by coastal factors such as salt spray.

Climate: The tropical climate ranges from mesic to wet. Moisture zones are seasonally mesic and moist mesic (Zones 4 and 5) on sites between the dry leeward and wet windward, and moderately wet to very wet windward sites (Zones 6 and 7) of the 7 Moisture Zones developed for the Hawaiian Islands by Price et al. (2007). The mesic forests have relatively even, moderate precipitation throughout the year or have a distinct, seasonally dry period and typically do not experience extended periods of drought like the dry forests and woodlands. Annual rainfall is 1200-2500 mm (47-98 inches) and falls largely from October to March. Annual rainfall in rainforests generally exceeds 2500 mm (98 inches).

Soil/substrate/hydrology: Substrates are highly variable ranging from gray acidic clays on older islands to thin organic mucks over lava flows and ash beds on Hawai`i. Soils are generally well-drained and include steep, rocky talus, shallow to deep soils over weathered rock and gravelly alluvium, rocky shallow organic muck.

Geographic Range: This lowland ruderal forest and woodland group is found in mesic to wet areas of Hawai`i, Moloka`i, Maui, Kaua`i and O`ahu.

Nations: US

States/Provinces:  HI




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNA

Greasons: No Data Available


Concept Lineage: G391 & G392 merged into G390.

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: >< Lowland Mesic Forest (Gagne and Cuddihy 1990)

Concept Author(s): K.A. Schulz, in Faber-Langendoen et al. (2016)

Author of Description: K.A. Schulz

Acknowledgements: No Data Available

Version Date: 05-26-16

  • Cuddihy, L. W., and C. P. Stone. 1990. Alteration of native Hawaiian vegetation: Effects of humans, their activities and introductions. University of Hawai''i Press for Cooperative National Parks Resource Studies Unit. Department of Botany, University of Hawai''i at Manoa, Honolulu. 138 pp.
  • Faber-Langendoen, D., J. Drake, S. Gawler, M. Hall, C. Josse, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, L. Sneddon, K. Schulz, J. Teague, M. Russo, K. Snow, and P. Comer, editors. 2010-2019a. Divisions, Macrogroups and Groups for the Revised U.S. National Vegetation Classification. NatureServe, Arlington, VA. plus appendices. [in preparation]
  • Gagne, W. C., and L. W. Cuddihy. 1990. Vegetation. Pages 45-114 in: W. L. Wagner, D. R. Herbst, and S. H. Sohmer, editors. Manual of the Flowering Plants of Hawaii. 2 volumes. University of Hawaii Press, Honolulu.
  • Price, J. P., S. M. Gon, III, J. D. Jacobi, and D. Matsuwaki. 2007. Mapping plant species ranges in the Hawaiian Islands: Developing a methodology and associated GIS layers. Hawai''i Cooperative Studies Unit. Technical Report HCSU-008. Pacific Aquaculture and Coastal Resources Center (PACRC), University of Hawai''i, Hilo. 58 pp., includes 16 figures and 6 tables.
  • Wagner, W. L., D. R. Herbst, and S. H. Sohmer. 1999. Manual of the flowering plants of Hawaii. Revised edition. Volumes 1 and 2. University of Hawaii Press and Bishop Museum Press, Honolulu. 1919 pp.