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G448 Caribbean Wet Montane Forest Group

Type Concept Sentence: Tall rainforests covered with epiphytes, growing above 800 m and up to nearly 2000 m elevation on tall mountains in Cuba, Jamaica, Puerto Rico and the Dominican Republic. They have a closed canopy and a second layer with abundant palms and tree ferns.


Common (Translated Scientific) Name: Caribbean Wet Montane Forest Group

Colloquial Name: Caribbean Wet Montane Forest

Hierarchy Level:  Group

Type Concept: This group includes tall rainforests growing above 800 m and up to nearly 2000 m elevation on tall mountains in Cuba, Jamaica, Puerto Rico and the Dominican Republic, where the moisture caused by almost daily presence of mist is enough to influence the structure and composition of the forest, characterized by bryophytes covering most tree trunks and branches, as well as being abundant in the ground layer. Bromeliads, ferns and orchids are also abundant as epiphytes and in some of these forests the presence of palms and tree ferns is common.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This group does not include the stunted or elfin cloud forest typical of the highest elevations, usually on windward slopes of mountains in the Greater Antilles, which often remain shrouded in fog. This group includes the so called Sierra Palm forest of Puerto Rico, characterized by stands dominated by the palm Prestoea acuminata var. montana.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Remnants of these evergreen tall forests have a closed or open canopy, 15-25 m high, consisting of microphylls and notophylls. When in good condition, the upper layer is closed and has a second layer with abundant palms, tree ferns and epiphytes, all of them rich in species. Lianas are rare but there are many climbing, scrambling and epiphytic shrubs. The shrub layer in general tends to be scattered, rarely reaching more that about 3 m, or in very humid occurrences the shrub layer forms an almost impenetrable bush tangled with pteridophytic lianas, as is the case in the high mountains of Pico Turquino in Cuba.

Floristics: As with the composition of other Caribbean groups, different islands have different species assemblages. In Puerto Rico, the following species are typical: Banara portoricensis, Brachionidium ciliolatum, Myrcia margarettiae (= Eugenia margarettiae), Gonocalyx concolor, Habenaria amalfitana (= Habenaria dussii), Ternstroemia luquillensis, and Ternstroemia subsessilis. In Cuba above 800 m elevation in the humid mountains of Sierra Maestra and Escambray typical species are Magnolia cubensis and various species of Ocotea such as the endemic Ocotea leucoxylon. These trees reach heights of about 25 m and below this a subcanopy can usually be characterized by Clusia tetrastigma, Gomidesia lindeniana, and several endemic species such as Fadyenia hookeri (= Garrya fadyenii) and Ossaea ottoschmidtii. At elevations above 1600 m with annual rainfall exceeding 3000 mm, the low canopy is characterized by various endemics such as Ageratina paucibracteata (= Eupatorium paucibracteatum), Myrsine microphylla, Nectandra reticularis, Persea anomala, Sapium maestrense, Symplocos leonis, Torralbasia cuneifolia, and several tree ferns. In Jamaica, in the upper reaches of the Blue Mountains, the dominant trees are Cyrilla racemiflora, Clusia havetioides, Eugenia marchiana, Ilex macfadyenii, Laplacea haematoxylon, Podocarpus urbanii, Rhamnus sphaerosperma, Solanum punctulatum, Turpinia occidentalis, and Viburnum villosum. Buchenavia tetraphylla (= Buchenavia capitata), Calyptronoma plumeriana (= Calyptromona dulcis), Cyathea arborea, Cyrilla racemiflora, Mora abbottii, Ormosia krugii, Prestoea acuminata var. montana (= Prestoea montana), and Sloanea berteriana are common in the montane humid broadleaf forests of Hispaniola.

Dynamics:  Landslides and hurricanes are the key triggers of dynamic processes of these forests. Substrate and topography and their interaction with the vegetation are the most important factors for the survival of these forests during hurricanes - probably the single most important natural trigger of the successional dynamic. Surviving trees have their roots securely anchored in the substrate. These factors are also critical for regulating surface runoff and maintaining the water balance under very humid conditions on exposed ridges and steep slopes. Forest recovery after disturbance is slow.

Environmental Description:  Forests within this group occur over 800 m and up to about 2000 m elevation on yellowish or red ferrallitic soils or clay-loam derived from limestones. In mountains exposed to higher precipitation, it is found as low as 450 m.

Climate: Due to elevational gradients and geographic position, the climate of the Caribbean wet montane forest is cooler and wetter than in the lowlands, with temperatures at high-elevation sites averaging 18.5°C or less and precipitation averaging greater than 3000 mm annually. These areas above 800 m experience frequent cloud cover and relative humidity near 100% due to passing low clouds and the upward transport of moisture from warm Caribbean waters.

Soil/substrate/hydrology: Forests growing on exposed areas are of smaller stature and very dense. Taller forests grow on protected areas on lower slopes to the leeward of ridges or spurs. With montane forests, one of the most critical climatic factors is the frequency and duration of the cloud cover; condensation can contribute 10% or more of the precipitation amount that these forests receive. In the Caribbean, the trade winds forming clouds have saline components which have an effect on the chemistry of the ecophysiology of these forests. Cloud cover causes less solar radiation, lower temperatures, decreased transpiration and lower photosynthetic rates, resulting in lower growth rates and lower nutrient-cycling rates. The efficiency shown by these forests in the use of nutrients is high though, which is important to avoid nutrient loss due to leaching.

Geographic Range: Forests within this group occur in the Luquillo Mountains of northeast Puerto Rico, Sierra Maestra and Sierra del Purial in south eastern Cuba, and the Escambray Mountains in the south-central region of Cuba, in the windward slopes of the Central and Northern Cordilleras and the Sierra de Neiba in the Dominican Republic, and the upper reaches of the Blue Mountains in Jamaica. Forests of this group might occur also on mountain slopes of several of the volcanic islands of the Lesser Antilles, but references have not been consulted.

Nations: CU,DO,JM,PR,XD

States/Provinces:  No Data Available



Confidence Level: Low - Poorly Documented

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Montane Forest Zone (Dansereau 1966)

Concept Author(s): C. Josse and D. Faber-Langendoen, in Faber-Langendoen et al. (2016)

Author of Description: C. Josse

Acknowledgements: No Data Available

Version Date: 02-04-16

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