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CEGL005297 Coleogyne ramosissima - Ephedra spp. Warm Desert Shrubland

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Blackbrush - Joint-fir species Warm Desert Shrubland

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This shrubland association is found at mid to upper elevations in the Mojave Desert and extends north and east into the Colorado Plateau and southern Great Basin in Arizona, California, Nevada, and southern Utah and often forms vast expanses across these environments. The vegetation is characterized by an open to patchy shrub canopy (9-29% cover) dominated or codominated by Coleogyne ramosissima with one or more species of Ephedra present to codominant. In the Mojave Desert and southern Great Basin, Ephedra nevadensis and Ephedra aspera are most common, whereas in the Colorado Plateau Ephedra torreyana is common with Ephedra cutleri less prevalent. Ephedra viridis typically occurs at higher elevation stands rangewide. Many other shrub associates are present across its range. If present, the herbaceous layer is sparse to open and composed of a mixture of grasses and forbs such as Achnatherum hymenoides, Achnatherum speciosum, Amsinckia spp., Chamaesyce serpyllifolia, Dasyochloa pulchella, Pleuraphis jamesii, Pleuraphis rigida, Poa secunda, Sphaeralcea grossulariifolia, Sporobolus cryptandrus, and exotic annuals Bromus rubens, Bromus tectorum, Erodium cicutarium, Salsola tragus, and Schismus barbatus. Stands can be seen on a variety of landforms. At lower elevations it occurs on ridgelines in transition areas with Larrea tridentata scrub and in gravelly and cobbly alluvial valleys, alluvial fans and hillslopes at mid to upper elevations, but includes eolian deposits/dunes, along drainage channels, basins, benches, mesas, terraces, plateaus, canyon slopes and rims, and ridges at 850 to 2133 m elevation. While predominantly found on flat or low-gradient settings, this association occasionally occurs on moderate to steep slopes. Soils are diverse, though primarily rapidly drained sandy loam or loam, and rarely finer-textured soils, and are derived from a variety of substrates. Dark cyanobacteria and mosses, if present, have sparse to low cover.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This association is largely known and described from National Park Service inventory efforts in the Colorado Plateau and Mojave Desert. The concept of this association is broad, including stands in the Mojave Desert, Great Basin and Colorado Plateau and is characterized by codominance of Coleogyne ramosissima with one of five Ephedra species. Within the range of this association some of the Ephedra spp. are more characteristic of the Colorado Plateau (Ephedra cutleri, Ephedra torreyana) or Mojave Desert and Great Basin (Ephedra aspera, Ephedra nevadensis) or cooler higher elevation sites rangewide (Ephedra viridis), but there is a mixing of Ephedra species in transition areas such as the Grand Canyon. Associated species are diverse, reflecting its wide range. Blackbrush shrublands cover vast areas in the Mojave Desert, Great Basin and Colorado Plateau and would greatly benefit from a rangewide analysis of plots to better clarify ecological community patterns.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: This blackbrush vegetation type often forms vast expanses across these environments and typically lacks trees and tall shrubs. The vegetation is characterized by an open to patchy shrub canopy (9-29% cover) dominated or codominated by Coleogyne ramosissima with one or more species of Ephedra present to codominant (Reid and Hall 2010, Evens et al. 2012, 2014). In the Mojave Desert and southern Great Basin, Ephedra nevadensis and Ephedra aspera (= Ephedra fasciculata) are most common, whereas in the Colorado Plateau Ephedra torreyana and Ephedra viridis are common with Ephedra cutleri less prevalent (Reid and Hall 2010). The widespread Ephedra viridis occurs at higher elevations rangewide (Peterson 1984a, Reid and Hall 2010, Evens et al. 2012, 2014). Other shrubs are often present, including Acacia greggii, Artemisia bigelovii, Atriplex canescens, Atriplex confertifolia, Atriplex obovata, Encelia frutescens, Ericameria cooperi, Ericameria nauseosa var. juncea (= Chrysothamnus nauseosus var. junceus), Ericameria teretifolia, Eriogonum corymbosum, Eriogonum fasciculatum, Glossopetalon spinescens, Grayia spinosa, Gutierrezia microcephala, Gutierrezia sarothrae, Krameria grayi, Krascheninnikovia lanata, Lycium andersonii, Opuntia polyacantha, Opuntia polyacantha var. erinacea (= Opuntia erinacea), Polygala acanthoclada, Psorothamnus arborescens, Psorothamnus fremontii, Rhus aromatica, Shepherdia rotundifolia, Tetradymia axillaris, Thamnosma montana, Viguiera parishii, Xylorhiza tortifolia, Yucca angustissima, Yucca baccata, and Yucca schidigera (Peterson 1984a, Reid and Hall 2010, Evens et al. 2012, 2014). Occasional emergent Yucca brevifolia trees are sometimes present in Mojave stands and Agave utahensis may occur in Colorado Plateau stands (Peterson 1984a, Reid and Hall 2010, Evens et al. 2012, 2014). If present, the herbaceous layer is sparse to open (to 14% cover) and composed of a mixture of grasses and low cover of forbs. Although usually sparse it may be diverse with species such as Abronia fragrans, Achnatherum hymenoides, Achnatherum speciosum, Amsinckia spp., Aristida purpurea, Bouteloua barbata, Calochortus kennedyi, Chamaesyce serpyllifolia, Dasyochloa pulchella, Elymus multisetus, Grindelia fastigiata, Lepidium montanum, Penstemon ambiguus, Plantago patagonica, Pleuraphis jamesii, Pleuraphis rigida, Poa secunda, Sphaeralcea grossulariifolia, Sporobolus contractus, Sporobolus cryptandrus, Tiquilia latior, Tridens muticus, Vulpia octoflora, and exotic annuals Bromus rubens, Bromus tectorum, Erodium cicutarium, Salsola tragus, and Schismus barbatus (Peterson 1984a, Reid and Hall 2010, Evens et al. 2012, 2014). Biological soil crusts, mostly in the form of dark cyanobacteria and mosses, may have up to 40% cover.

Dynamics:  No Data Available

Environmental Description:  This shrubland association is found at mid to upper elevations in the Mojave Desert and extends north and east into the Colorado Plateau and southern Great Basin in Arizona, California, Nevada and southern Utah and often forms vast expanses across these environments. It can be seen on a variety of landforms. At lower elevations it occurs on ridgelines in transition areas with Larrea tridentata scrub and in gravelly and cobbly alluvial valleys, alluvial fans and hillslopes at mid to upper elevations, but includes eolian deposits/dunes, along drainage channels, basins, benches, mesas, terraces, plateaus, canyon slopes and rims, and ridges at 850 to 2133 m elevation (Peterson 1984a, Reid and Hall 2010, Evens et al. 2012, 2014). While predominantly found on flat or low-gradient settings (0-15°), this association occasionally occurs on moderate to steep slopes (to 40°) (Reid and Hall 2010, Evens et al. 2012, 2014). Soils are diverse, though primarily rapidly drained sand or loam, and less frequently finer-textured soils such as moderately well-drained to somewhat poorly drained silt loams, silty clay loams, clay loams, silts, or clays (Reid and Hall 2010, Evens et al. 2012, 2014). Sand or bare soil covers most of the unvegetated surface, with low to moderate cover by small rocks, large rocks, litter, and bedrock. Soils are derived from a variety of substrates such as alluvium, eolian deposits, dolostone (dolomite), granodiorite, quartz monzonite, limestone, rhyolite, sandstones of the Navajo, Kayenta, Entrada, or White Rim formations and shales of the Carmel, Halgaito, Moenkopi, Morrison, or Shinarump formations (Peterson 1984a, Reid and Hall 2010, Evens et al. 2012, 2014). Dark cyanobacteria and mosses, if present, have sparse to low cover.

Geographic Range: This shrubland association is found at mid to upper elevations in the Mojave Desert and southern Great Basin in Arizona, California and Nevada. It was sampled in Death Valley National Park, Glen Canyon National Recreation Area, Grand Canyon National Park, Mojave National Preserve and Grand Canyon-Parashant National Monument.

Nations: US

States/Provinces:  AZ, CA, NV, UT




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Coleogyne ramosissima - Ephedra nevadensis (Sawyer et al. 2009) [33.020.03]
= Coleogyne ramosissima - Ephedra nevadensis Association (Evens et al. 2012)
= Coleogyne ramosissima - Ephedra spp. Shrubland (Reid and Hall 2010)
= Coleogyne ramosissima - Ephedra spp. Warm Desert Shrubland (Reid and Hall 2010)
= Coleogyne ramosissima-Ephedra nevadensis Association (Evens et al. 2014)
= Coleogyne ramosissima-Ephedra nevadensis Association (Peterson 1984a)
= Coleogyne ramosissima-Ephedra nevadensis Association (Keeler-Wolf and Thomas 2000)
< Coleogyne ramosissima Shrubland Alliance (Keeler-Wolf and Thomas 2000)
< Black Brush Series (Sawyer and Keeler-Wolf 2005)

Concept Author(s): M.S. Reid and M.E. Hall (2010)

Author of Description: R. Meszaros and K.A. Schulz

Acknowledgements: No Data Available

Version Date: 11-30-16

  • Evens, J. M., D. Roach-McIntosh, and D. Stout. 2012. Vegetation descriptions for Joshua Tree National Park. Unpublished report submitted to USDI, National Park Service, Mojave Desert Inventory and Monitoring Network. California Native Plant Society, Sacramento, CA.
  • Evens, J. M., K. Sikes, D. Hastings, and J. Ratchford. 2014. Vegetation alliance descriptions for Lake Mead National Recreation Area, Death Valley National Park and Mojave National Preserve. Unpublished report submitted to USDI National Park Service, Mojave Desert Network Inventory and Monitoring Program. California Native Plant Society, Sacramento, CA.
  • Kearsley, M. J. C., K. Green, M. Tukman, M. Reid, M. Hall, T. J. Ayers, and K. Christie. 2015. Grand Canyon National Park-Grand Canyon / Parashant National Monument vegetation classification and mapping project. Natural Resource Report NPS/GRCA/NRR--2015/913. National Park Service, Fort Collins, CO. 75 pp. plus appendices.
  • Keeler-Wolf, T., and K. Thomas. 2000. Draft descriptions of vegetation alliances for the Mojave Ecosystem Mapping project. California Natural Diversity Database, California Department of Fish and Game, Sacramento.
  • Peterson, P. M. 1984a. Flora and physiognomy of the Cottonwood Mountains, Death Valley National Monument, California. University of Nevada Cooperative National Park Resources Studies Unit Report CPSU/UNLV 022/06. Las Vegas, NV.
  • Reid, M. S., and M. E. Hall. 2010. Vegetation classification of Grand Canyon National Park. Draft report submitted to National Park Service, Fort Collins, CO.
  • Sawyer, J. O., T. Keeler-Wolf, and J. Evens. 2009. A manual of California vegetation. Second edition. California Native Plant Society, Sacramento CA. 1300 pp.
  • Sawyer, J. O., and T. Keeler-Wolf. 2005. A manual of California vegetation. II. Draft manuscript. California Native Plant Society, Sacramento, CA.
  • Tendick, A., J. Spence, M. Reid, K. Shulz, G. Kittel, K. Green, A. Wight, and G. Wakefield. 2017. Vegetation classification and mapping project report, Glen Canyon National Recreation Area and Rainbow Bridge National Monument. Natural Resource Report NPS/NCPN/NRR—2017/1500. National Park Service, Fort Collins, CO. 1464 pp.
  • Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.