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G213 Pinus ponderosa var. ponderosa Central Rocky Mountain Open Woodland Group

Type Concept Sentence: This inland Pacific Northwest woodland and savanna group is found in the foothills of the central Rocky Mountains in the Columbia Plateau region and west along the foothills of the Modoc Plateau and eastern Cascades into southern interior British Columbia, occurring at the lower treeline/ecotone between grasslands or shrublands and more mesic coniferous forests, typically on warm, dry, exposed sites. This group includes two physiognomic phases: true woodlands of Pinus ponderosa with shrubby or grassy understories, and "wooded steppes" with widely spaced, scattered Pinus ponderosa trees over generally shrubby but sparse understories.


Common (Translated Scientific) Name: Ponderosa Pine Central Rocky Mountain Open Woodland Group

Colloquial Name: Central Rocky Mountain Ponderosa Pine Open Woodland

Hierarchy Level:  Group

Type Concept: This inland Pacific Northwest group occurs in the foothills of the central Rocky Mountains in the Columbia Plateau region and west along the foothills of the Modoc Plateau and eastern Cascades into southern interior British Columbia. It also occurs east across Idaho into the eastern foothills of the Montana Rockies. This group includes two physiognomic phases: true woodlands of Pinus ponderosa with shrubby or grassy understories, and "wooded steppes" with widely spaced, scattered Pinus ponderosa trees over generally shrubby but sparse understories. The former are generally fire-maintained, while the later are often too dry and with widely spaced vegetation to carry fire. Pinus ponderosa var. ponderosa is the predominant conifer; Pseudotsuga menziesii or Pinus flexilis may be present in the tree canopy but are usually absent. The understory can be shrubby, with Amelanchier alnifolia, Arctostaphylos patula, Arctostaphylos uva-ursi, Artemisia tridentata, Cercocarpus ledifolius, Physocarpus malvaceus, Purshia tridentata, Rosa spp., Symphoricarpos albus, or Symphoricarpos oreophilus as common species. In transition areas with sagebrush steppe, Artemisia tridentata ssp. tridentata, Artemisia tridentata ssp. wyomingensis, Artemisia tripartita, and Purshia tridentata may be common in fire-protected sites such as rocky areas. Deciduous shrubs, such as Physocarpus malvaceus, Spiraea betulifolia, or Symphoricarpos albus, can be abundant in more northerly sites or more moist climates. Herbaceous vegetation in the true savanna occurrences is predominantly fire-resistant grasses and forbs that resprout following surface fires; shrubs, understory trees and downed logs are uncommon. These more open stands support grasses such as Achnatherum spp., dry Carex species (Carex inops), Elymus elymoides, Festuca campestris, Festuca idahoensis, Hesperostipa spp., or Pseudoroegneria spicata. These woodlands and savannas occur at the lower treeline/ecotone between grasslands or shrublands and more mesic coniferous forests, typically on warm, dry, exposed sites. These interior Pacific Northwest woodlands receive winter and spring rains, and thus have a greater spring "green-up" than the drier woodlands in the Central Rockies. However, sites are often too droughty to support a closed tree canopy. Elevations range from less than 500 m in British Columbia to 1600 m in the central Idaho mountains. Occurrences are found on all slopes and aspects; however, moderately steep to very steep slopes or ridgetops and plateaus are most common. This group generally occurs on most geological substrates from weathered rock to glacial deposits to eolian deposits. Characteristic soil features include good aeration and drainage, coarse textures, circumneutral to slightly acidic pH, an abundance of mineral material, and periods of drought during the growing season. Some occurrences may occur as edaphic climax communities on very skeletal, infertile and/or excessively drained soils, such as pumice, cinder or lava fields, and scree slopes. Surface textures are highly variable in this group, ranging from sand to loam and silt loam. Exposed rock and bare soil consistently occur to some degree in all the associations. The more mesic portions of this group may include Calamagrostis rubescens or Carex geyeri, species more typical of ~Central Rocky Mountain Douglas-fir - Pine Forest Group (G210)$$. Mixed fire regimes and surface fires of variable return intervals maintain these woodlands, typically with a shrub-dominated or patchy shrub layer, depending on climate, degree of soil development, and understory density. Historically, many of these woodlands and savannas lacked the shrub component as a result of 3- to 7-year fire-return intervals.

Diagnostic Characteristics: Pinus ponderosa var. ponderosa woodlands, savannas and "wooded steppes" in the central Rocky Mountains and eastern Cascades, lacking other conifers and oaks. Generally other floristic components are Central Rockies or somewhat similar to the northern Basin and Range region.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This includes the northern race of Interior Ponderosa Pine old-growth (USFS Region 6, USFS Region 1). The FEIS site describes different varieties of Pinus ponderosa and associated species. This group of the Central Rockies is primarily Pinus ponderosa var. ponderosa (Habeck 1992). Johansen and Latta (2003) have mapped the distribution of two varieties (Pinus ponderosa var. scopulorum and Pinus ponderosa var. ponderosa) using mitochondrial DNA. They hybridize along the Continental Divide in Montana backing up the FEIS information. ~Southern Rocky Mountain Ponderosa Pine Forest & Woodland Group (G228)$$ and ~Southern Rocky Mountain Ponderosa Pine Open Woodland Group (G229)$$ mostly contain Pinus ponderosa var. scopulorum, Pinus ponderosa var. brachyptera, and Pinus arizonica var. arizonica. The transition from this group to G228 is now defined to occur in the montane zones of the Bighorns (USFS section M331B) and Laramie Range (USFS section M331I) and to the east and south of these mountains. The Southern Rocky Mountain group will also occur in other isolated mountain ranges of central Wyoming, but not in eastern Wyoming. It does not occur farther north than Wyoming; all western Montana ponderosa pine woodlands are placed into this ~Central Rocky Mountain Ponderosa Pine Open Woodland Group (G213)$$. Ponderosa woodlands and "steppes" in eastern Wyoming, eastern and central Montana, including the Missouri River Breaks, are now included in ~Black Hills-Northwestern Great Plains Ponderosa Pine Forest & Woodland Group (G216)$$, which is predominantly Pinus ponderosa var. scopulorum. Ponderosa pine woodlands found in the Great Plains do show some floristic similarities to these found within the forested mountains of the Central Rockies, but typically have herbaceous floristics related to the Great Plains "mixedgrass." South of the Modoc Plateau in California, Pinus ponderosa forests and woodlands are included in ~Californian Montane Conifer Forest & Woodland Group (G344)$$.

Woodlands dominated by Pseudotsuga menziesii found in breaks along rivers and on escarpments in central and eastern Montana and Wyoming are not included in this group, they are placed with ~Middle Rocky Mountain Montane Douglas-fir Forest & Woodland Group (G215)$$ or ~Central Rocky Mountain Douglas-fir - Pine Forest Group (G210)$$. Hot, dry Douglas-fir types with grass are also not included here, although they are floristically similar. In southern interior British Columbia, Pseudotsuga menziesii or Pinus flexilis may form woodlands or fire-maintained savannas with and without Pinus ponderosa var. ponderosa at the lower treeline transition into grassland or shrub-steppe. In interior British Columbia, Pseudotsuga menziesii is the characteristic canopy dominant. These Douglas-fir and limber pine associations are currently placed into other groups.

A meeting of Pacific Northwest ecologists for Landfire concluded that the "true savannas" of high-frequency / low-intensity fires and grassy understories are now rare. See Dynamics section for more information.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: These are conifer-dominated woodlands, forests, and savannas, with grassy or shrubby understories. Shrubs can be broad-leaved deciduous or microphyllous evergreen (sagebrush), while the graminoids are primarily bunch grasses, along with rhizomatous grasses. Understories are generally low to moderate in cover, especially in the most droughty and rocky sites. In some cases due to a climate-edaphic interaction, the structure is that of widely scattered trees over "shrub-steppe" of sage, bitterbrush, or sparsely distributed grasses.

Floristics: This group includes two physiognomic phases: true woodlands of Pinus ponderosa with shrubby or grassy understories, and "wooded steppes" with widely spaced, scattered Pinus ponderosa trees over generally shrubby but sparse understories. The former are generally fire-maintained, while the later are often too dry and with widely spaced vegetation to carry fire. Pinus ponderosa var. ponderosa is the predominant conifer; Pinus flexilis or Pseudotsuga menziesii may be present in the tree canopy but are usually absent. The understory can be shrubby, with Amelanchier alnifolia, Arctostaphylos patula, Arctostaphylos uva-ursi, Ceanothus velutinus, Cercocarpus ledifolius, Physocarpus malvaceus, Purshia tridentata, Rosa spp., Symphoricarpos albus or Symphoricarpos oreophilus, and Vaccinium cespitosum as common species. In transition areas with big sagebrush steppe systems, Artemisia arbuscula, Artemisia tridentata ssp. tridentata, Artemisia tridentata ssp. wyomingensis, Artemisia tripartita, Artemisia tridentata ssp. wyomingensis, Artemisia tripartita, and Purshia tridentata may be common in fire-protected sites such as rocky areas. Deciduous shrubs, such as Physocarpus malvaceus, Spiraea betulifolia, or Symphoricarpos albus, can be abundant in more northerly sites or more moist climates. Understory vegetation in the true savanna occurrences is predominantly fire-resistant grasses and forbs that resprout following surface fires; shrubs, understory trees and downed logs are uncommon. These more open stands support grasses such as Achnatherum spp., dry Carex species (Carex inops), Festuca campestris, Festuca idahoensis, Hesperostipa spp., or Pseudoroegneria spicata. The more mesic portions of this group may include Calamagrostis rubescens or Carex geyeri, species more typical of ~Central Rocky Mountain Douglas-fir - Pine Forest Group (G210)$$.

Dynamics:  Pinus ponderosa is a drought-resistant, shade-intolerant conifer which usually occurs at lower treeline in the major ranges of the western United States. Historically, surface fires and drought were influential in maintaining open-canopy conditions in these woodlands. With settlement and subsequent fire suppression, occurrences have become denser. Presently, some occurrences contain understories of more shade-tolerant species, such as Pseudotsuga menziesii and/or Abies spp., as well as younger cohorts of Pinus ponderosa. These altered structures have affected fuel loads and fire regimes. Presettlement fire regimes were primarily frequent (5- to 15-year return intervals), low-intensity surface fires triggered by lightning strikes or deliberately set fires by Native Americans. With fire suppression and increased fuel loads, fire regimes are now less frequent and often become intense crown fires, which can kill mature Pinus ponderosa (Reid et al. 1999). Establishment is erratic and believed to be linked to periods of adequate soil moisture and good seed crops as well as fire frequencies, which allow seedlings to reach sapling size. Longer fire-return intervals have resulted in many occurrences having dense subcanopies of overstocked and unhealthy young Pinus ponderosa, along with Pseudotsuga menziesii on moist sites (Reid et al. 1999).

Additional associations included in this group are subject to periodic drought that limits tree establishment. This climate-edaphic interaction results in widely scattered trees over "shrub-steppe" of sagebrush, bitterbrush, or sparsely distributed grasses. Tree growth is likely episodic, with regeneration episodes in years with available moisture. Tree density is limited in some areas by available growing space due to rocky conditions of the site. Hence the tree canopy in these associations will never reach woodland density or close due to the interaction of climate and edaphic factors, even in the absence of fire. They burn occasionally, but the vegetation is sparse enough that fires are typically not carried through the stand. Fire frequency is speculated to be 30-50 years. Some stands also occur on areas of sand dunes, scablands, and pumice where the edaphic conditions limit tree abundance.

A meeting of Pacific Northwest ecologists for Landfire concluded that the "true savannas" of high-frequency / low-intensity fires and grassy understories are now rare. Most areas that may have been savanna in the past are now more nearly closed-canopy woodlands/forests. Conclusion was that these true savannas should be included with this woodland group, along with the climatically-edaphically controlled "wooded steppes" which are also in this group. The wooded steppes included here are not fire-maintained; they occur on sites too droughty to support a closed tree canopy. They do burn with a high-frequency / low-intensity regime, but fire is not carried because of the sparse vegetation of the edaphically constrained sites (rock outcrops, dunes, super-dry, sparse trees over shrubs and sometimes grasses but widely spaced). Louisa Evers (pers. comm. 2006) notes that she has not found any evidence that ponderosa pine savanna existed historically in north-central and central Oregon. In north-central Oregon, the savanna would have been oak or pine-oak. In central Oregon, it may well have been western juniper. Condition surveys of the Cascades Forest Reserve and General Land Office survey notes suggest that ponderosa pine formed a woodland with grassy understories, but still was often referred to as open-parklike. Conversely pine-oak and Douglas-fir-oak savannas appeared to have once been quite common in the Willamette Valley.

Environmental Description:  This group within the interior Pacific Northwest region occurs at the lower treeline/ecotone between grasslands or shrublands and more mesic coniferous forests typically in warm, dry, exposed sites at elevations ranging from 500-1600 m (1600-5248 feet). These interior Pacific Northwest woodlands receive winter and spring rains, and thus have a greater spring "green-up" than the drier woodlands in the Central Rockies. However, these sites are often too droughty to support a closed tree canopy. They can occur on all slopes and aspects; however, they commonly occur on moderately steep to very steep slopes or ridgetops and plateaus. Substrates include glacial till, glacio-fluvial sand and gravel, dunes, basaltic rubble and scablands, colluvium, or deep loess or volcanic ash-derived soils, all with characteristic features of good aeration and drainage, coarse textures, circumneutral to slightly acidic pH, an abundance of mineral material, rockiness, and periods of drought during the growing season. In the Oregon "pumice zone" this group occurs as matrix-forming, extensive woodlands on rolling pumice plateaus and other volcanic deposits.

Climate: This group within the interior Pacific Northwest region occurs at the lower treeline/ecotone between grasslands or shrublands and more mesic coniferous forests typically in warm, dry, exposed sites at elevations ranging from 500-1600 m (1600-5248 feet). These interior Pacific Northwest woodlands receive winter and spring rains, and thus have a greater spring "green-up" than the drier woodlands in the Central Rockies.

Soil/substrate/hydrology: These sites are often too droughty to support a closed tree canopy. They can occur on all slopes and aspects; however, they commonly occur on moderately steep to very steep slopes or ridgetops and plateaus. Substrates include glacial till, glacio-fluvial sand and gravel, dunes, basaltic rubble and scablands, colluvium, or deep loess or volcanic ash-derived soils, all with characteristic features of good aeration and drainage, coarse textures, circumneutral to slightly acidic pH, an abundance of mineral material, rockiness, and periods of drought during the growing season. In the Oregon "pumice zone" this group occurs as matrix-forming, extensive woodlands on rolling pumice plateaus and other volcanic deposits.

Geographic Range: This group is found in the Fraser River drainage of southern British Columbia south along the Cascades and central Rocky Mountains of Washington, Oregon and the Modoc Plateau of northeastern California. In the northeastern part of its range, it extends across the central Rocky Mountains west of the Continental Divide into northwestern Montana, south to the Snake River Plain in Idaho, and east into the foothills of western Montana (but not into central or eastern Montana). In Oregon, it is most common in south-central Oregon, in lands managed by the Lakeview District of the BLM, and by the adjacent Fremont and Deschutes national forests. It also occurs on the marginal lands coming south out of the Blue Mountains, on the edge of the northern Basin and Range.

Nations: CA,US

States/Provinces:  AB, BC, CA, CO, ID, MT, ND, NM, NV?, OR, SD, UT, WA, WY




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: No Data Available

Concept Author(s): R.F. Daubenmire and J.B. Daubenmire (1968)

Author of Description: M.S. Reid, K.A. Schulz and M. Manning

Acknowledgements: M. Manning

Version Date: 11-09-15

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