Print Report

G300 Atriplex confertifolia - Atriplex canescens - Grayia spinosa Scrub Group

Type Concept Sentence: This widespread semi-arid scrub group is found in basins, plains, alluvial flats and slopes in the intermountain western U.S. and western Great Plains and is characterized by a typically open to moderately dense shrub layer dominated or codominated by Atriplex canescens, Atriplex confertifolia, Atriplex cuneata, Atriplex hymenelytra, Atriplex obovata, Atriplex polycarpa, Grayia spinosa, and/or Picrothamnus desertorum often with other shrubs present to codominant.


Common (Translated Scientific) Name: Shadscale Saltbush - Fourwing Saltbush - Spiny Hopsage Scrub Group

Colloquial Name: Intermountain Shadscale - Saltbush Scrub

Hierarchy Level:  Group

Type Concept: This extensive group includes open-canopied shrublands of typically saline basins, alluvial slopes and plains across the intermountain western U.S. and western Great Plains. The vegetation is characterized by a typically open to moderately dense shrubland composed of one or more Atriplex species, such as Atriplex canescens, Atriplex confertifolia, Atriplex cuneata, Atriplex hymenelytra, Atriplex obovata, or Atriplex polycarpa. Other shrubs present to codominant may include Artemisia tridentata ssp. wyomingensis, Chrysothamnus viscidiflorus, Ericameria nauseosa, Ephedra nevadensis, Krascheninnikovia lanata, Grayia spinosa, Lycium spp., Picrothamnus desertorum, or Tetradymia spp. Northern occurrences lack Atriplex species and are typically dominated by Grayia spinosa. In Wyoming, occurrences are typically a mix of Artemisia tridentata ssp. wyomingensis, Atriplex confertifolia, Grayia spinosa, Krascheninnikovia lanata, Sarcobatus vermiculatus, and various Ericameria or Chrysothamnus species. Some places are a mix of Atriplex confertifolia and Artemisia tridentata ssp. wyomingensis. In the Great Basin, Sarcobatus vermiculatus is generally absent but, if present, does not codominate. The herbaceous layer varies from sparse to moderately dense and is dominated by perennial graminoids such as Achnatherum hymenoides, Bouteloua gracilis, Elymus lanceolatus ssp. lanceolatus, Pascopyrum smithii, Pleuraphis jamesii, Pleuraphis rigida, Poa secunda, or Sporobolus airoides. Various forbs are also present. Substrates are often saline and calcareous, medium- to fine-textured, alkaline soils, but include some coarser-textured soils.

Diagnostic Characteristics: The vegetation is characterized by a typically open to moderately dense shrubland composed of one or more shrub species, such as Atriplex canescens, Atriplex confertifolia, Atriplex cuneata, Atriplex hymenelytra, Atriplex obovata, Atriplex polycarpa, Grayia spinosa, and/or Picrothamnus desertorum.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Some Grayia spinosa-dominated stands tends to occur on coppice dunes that have a silty component to them. If they occur on deep sand or dunes, then consider a dune group. Scrub stands dominated by Picrothamnus desertorum are placed in this group because Picrothamnus desertorum is frequently present to codominant with the widespread shrub Atriplex confertifolia in the Great Basin (Mozingo 1987). However, Picrothamnus desertorum is often short (10-25 cm tall) so could also be classified as ~Intermountain Dwarf Saltbush - Sagebrush Scrub Group (G301)$$.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: This group is a facultatively deciduous, extremely xeromorphic, subdesert short shrubland with a typically open shrub canopy and sparse to moderately dense herbaceous layer dominated by perennial graminoids.

Floristics: This widespread cool semi-desert scrub group is highly variable and ranges from almost pure occurrences of a single species to fairly complex mixtures. The characteristic mix of low shrubs and grasses is sparse, with large open spaces between the plants (Blaisdell and Holmgren 1984). Occurrences have a sparse to moderately dense cover of woody species that is often dominated by one of several shrubs such as Atriplex canescens (also codominated by Artemisia tridentata, Ephedra viridis, or Krascheninnikovia lanata), Atriplex confertifolia (also codominated by several species of Lycium and Ephedra, Picrothamnus desertorum, or Sarcobatus vermiculatus), Atriplex cuneata, Atriplex obovata, Atriplex hymenelytra, Atriplex polycarpa, Grayia spinosa, or Picrothamnus desertorum (= Artemisia spinescens). Other shrubs may be present, especially in transition areas with desert or montane scrub. Species include Acacia greggii, Artemisia frigida, Chrysothamnus spp., Encelia frutescens, Ericameria nauseosa, Ephedra nevadensis, Frankenia salina, Larrea tridentata, Lycium andersonii, Lycium pallidum, Parthenium confertum, Psorothamnus polydenius, Purshia stansburiana, Suaeda spp., Tetradymia glabrata, Tetradymia spinosa, Tiquilia latior, and Yucca glauca. Dwarf-shrubs include Gutierrezia sarothrae and several species of Eriogonum, but generally do not form a distinct layer as the main shrub layer is often >0.5 m tall.

Warm-season medium-tall and short perennial grasses dominate in the sparse to moderately dense graminoid layer. The species present depend on the geographic range of the grasses, soil alkalinity/salinity, and past land use. Species may include Achnatherum hymenoides, Bouteloua gracilis, Distichlis spicata, Elymus elymoides, Hesperostipa comata, Leymus ambiguus, Leymus salinus, Muhlenbergia torreyi, Pascopyrum smithii, Pleuraphis jamesii, Poa secunda, Pseudoroegneria spicata, Sporobolus airoides, and Sporobolus cryptandrus. A number of annual species may also grow in association with the shrubs and grasses, although they are usually rare and confined to areas of recent disturbance (Blaisdell and Holmgren 1984). Forb cover is generally sparse. Perennial forbs that might occur include Chaetopappa ericoides, Descurainia spp., Mentzelia spp., Sphaeralcea coccinea, and Xylorhiza venusta. Annual natives include Monolepis nuttalliana, Plantago spp., or Vulpia octoflora. Associated halophytic annuals include Salicornia bigelovii, Salicornia rubra, and Suaeda species. Exotic annuals that may occur include Salsola kali, Bromus rubens, and Bromus tectorum. Cacti such as species of Opuntia and Echinocereus may be present in some occurrences. Trees are not usually present but some scattered Juniperus spp. may be found.

Dynamics:  West (1982) stated that "salt desert shrub vegetation occurs mostly in two kinds of situations that promote soil salinity, alkalinity, or both. These are either at the bottom of drainages in enclosed basins or where marine shales outcrop." However, salt-desert shrub vegetation may be an indication of climatically dry as well as physiologically dry soils (Blaisdell and Holmgren 1984). Not all salt-desert shrub soils are salty, and their hydrologic characteristics may often be responsible for the associated vegetation (Naphan 1966). Species of the salt-desert shrub complex have different degrees of tolerance to salinity and aridity, and they tend to sort themselves out along a moisture/salinity gradient (West 1982). Species and communities are apparently sorted out along physical, chemical, moisture, and topographic gradients through complex relations that are not understood and are in need of further study (Blaisdell and Holmgren 1984).

The winter months are a good time for soil moisture accumulation and storage within stands in this group. There is generally at least one good snowstorm per season that will provide sufficient moisture to the vegetation. The winter moisture accumulation amounts will affect spring plant growth. Plants may grow as little as a few inches to 1 m. Unless more rains come in the spring, the soil moisture will be depleted in a few weeks, growth will slow and ultimately cease, and the perennial plants will assume their various forms of dormancy (Blaisdell and Holmgren 1984). If effective rain comes later in the warm season, some of the species will renew their growth from the stage at which it had stopped. Others, having died back, will start over as if emerging from winter dormancy (Blaisdell and Holmgren 1984). Atriplex confertifolia shrubs often develop large leaves in the spring, which increase the rate of photosynthesis. As soil moisture decreases, the leaves are lost, and the plant takes on a dead appearance. During late fall, very small overwintering leaves appear which provide some photosynthetic capability through the remainder of the year. Other communities are maintained by intra- or inter-annual cycles of flooding followed by extended drought, which favor accumulation of transported salts. The moisture supporting these intermittently flooded wetlands is usually derived off-site, and they are dependent upon natural watershed function for persistence (Reid et al. 1999).

In summary, desert communities of perennial plants are dynamic and changing. The composition within this group may change dramatically and may be both cyclic and unidirectional. Superimposed on the compositional change is great variation from year to year in growth of all the vegetation, the sum of varying growth responses of individual species to specific conditions of different years (Blaisdell and Holmgren 1984). Desert plants grow when temperature is satisfactory, but only if soil moisture is available at the same time. Because the amount of moisture is variable from year to year and because different species flourish under different seasons of soil moisture, seldom do all components of the vegetation thrive in the same year (Blaisdell and Holmgren 1984).

Environmental Description:  This salt-desert shrubland group is matrix-forming in the Intermountain West. This group composes arid to semi-arid shrublands on lowland and upland sites usually at elevations between 1520 and 2200 m (4987-7218 feet). Sites can be found on all aspects and include valley bottoms, alluvial and alkaline flats, mesas and plateaus, playas, drainage terraces, washes and interdune basins, bluffs, and gentle to moderately steep sandy or rocky slopes. Slopes are typically gentle to moderately steep but are sometimes unstable and prone to surface movement. Many areas within this group are degraded due to erosion and may resemble "badlands." Soil surface is often very barren and interspaces between the characteristic plant clusters are commonly covered by a microphytic crust (West 1982).

Climate: This is typically a vegetation group of extreme climatic conditions, with warm to hot summers and freezing winters. Annual precipitation ranges from approximately 13-33 cm. In much of the group, the period of greatest moisture will be mid to late summer, although in the more northern areas, a moist period is to be expected in the cold part of the year. However, plotted seasonality of occurrence is probably of less importance on this desert vegetation than in other types because desert precipitation comes with an extreme irregularity that does not appear in graphs of long-term seasonal or monthly averages (Blaisdell and Holmgren 1984). Soil/substrate/hydrology: Soils are shallow to moderately deep, poorly developed, and a product of an arid climate and little precipitation. Soils are often alkaline or saline. Vegetation within this group is tolerant of these soil conditions but not restricted to it. The shallow soils of much of the area are poorly developed Entisols. Vegetation within this group can occur on level pediment remnants where coarse-textured and well-developed soil profiles have been derived from sandstone gravel and are alkaline, or on Mancos shale badlands, where soil profiles are typically fine-textured and non-alkaline throughout (West and Ibrahim 1968). They can also occur in alluvial basins where parent materials from the other habitats have been deposited over Mancos shale and the soils are heavy-textured and saline-alkaline throughout the profile (West and Ibrahim 1968).

Geographic Range: The distribution of this widespread group centers in the Intermountain West of the U.S., and extends in limited distribution across northern New Mexico into the southern Great Plains. In Wyoming, this group occurs in the Great Divide and Bighorn basins.

Nations: MX,US

States/Provinces:  AZ, CA, CO, ID, KS, MT, MXCHH, MXCOA, NM, NV, OR, TX, UT, WA, WY




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: < Biotic Matrix of the Shadscale and Associated Communities (Fautin 1946)
= Intermountain salt desert shrublands (West 1983b)
< Salt Desert Shrub (414) (Shiflet 1994)

Concept Author(s): N.E. West (1983b)

Author of Description: K.A. Schulz

Acknowledgements: No Data Available

Version Date: 11-06-15

  • Barbour, M. G., and J. Major, editors. 1988. Terrestrial vegetation of California: New expanded edition. California Native Plant Society, Special Publication 9, Sacramento. 1030 pp.
  • Blaisdell, J. P., and R. C. Holmgren. 1984. Managing intermountain rangelands-salt-desert shrub ranges. General Technical Report INT-163. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 52 pp.
  • Branson, F. A., R. F. Miller, and I. S. McQueen. 1967. Geographic distribution and factors affecting the distribution of salt desert shrubs in the United States. Journal of Range Management 29(5):287-296.
  • Branson, F. A., R. F. Miller, and I. S. McQueen. 1976. Moisture relationships in twelve northern desert shrub communities near Grand Junction, Colorado. Ecology 57:1104-1124.
  • Brown, D. E., C. H. Lowe, and C. P. Pase. 1979. A digitized classification system for the biotic communities of North America with community (series) and association examples for the Southwest. Journal of the Arizona-Nevada Academy of Science 14:1-16.
  • Brown, D. E., editor. 1982a. Biotic communities of the American Southwest-United States and Mexico. Desert Plants Special Issue 4(1-4):1-342.
  • Campbell, V. O. 1977. Certain edaphic and biotic factors affecting vegetation in the shadscale community of the Kaiparowitz area. Unpublished thesis, Brigham Young University, Provo, UT. 59 pp.
  • Faber-Langendoen, D., J. Drake, S. Gawler, M. Hall, C. Josse, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, L. Sneddon, K. Schulz, J. Teague, M. Russo, K. Snow, and P. Comer, editors. 2010-2019a. Divisions, Macrogroups and Groups for the Revised U.S. National Vegetation Classification. NatureServe, Arlington, VA. plus appendices. [in preparation]
  • Fautin, R. W. 1946. Biotic communities of the Northern Desert Shrub Biome in western Utah. Ecological Monographs 16(4):251-310.
  • Francis, R. E. 1986. Phyto-edaphic communities of the Upper Rio Puerco Watershed, New Mexico. Research Paper RM-272. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 73 pp.
  • Holland, V. L., and D. J. Keil. 1995. California vegetation. Kendall/Hunt Publishing Company, Dubuque, IA. 516 pp.
  • Knight, D. H. 1994. Mountains and plains: Ecology of Wyoming landscapes. Yale University Press, New Haven, MA. 338 pp.
  • Knight, D. H., G. P. Jones, Y. Akashi, and R. W. Myers. 1987. Vegetation ecology in the Bighorn Canyon National Recreation Area. Unpublished report prepared for the USDI National Park Service and University of Wyoming-National Park Service Research.
  • Mozingo, H. 1987. Shrubs of the Great Basin: A natural history. University of Nevada Press, Las Vegas. 342 pp.
  • Naphan E. A. 1966. Soils of the salt desert shrub areas and their productive capabilities. Pages 44-68 in: Proceeding: Salt desert shrub symposium. USDI, Bureau of Land Management. Cedar City, UT.
  • Reid, M. S., K. A. Schulz, P. J. Comer, M. H. Schindel, D. R. Culver, D. A. Sarr, and M. C. Damm. 1999. An alliance level classification of vegetation of the coterminous western United States. Unpublished final report to the University of Idaho Cooperative Fish and Wildlife Research Unit and National Gap Analysis Program, in fulfillment of Cooperative Agreement 1434-HQ-97-AG-01779. The Nature Conservancy, Western Conservation Science Department, Boulder, CO.
  • Shiflet, T. N., editor. 1994. Rangeland cover types of the United States. Society for Range Management. Denver, CO. 152 pp.
  • Stout, D., J. Buck-Diaz, S. Taylor, and J. M. Evens. 2013. Vegetation mapping and accuracy assessment report for Carrizo Plain National Monument. California Native Plant Society, Vegetation Program, Sacramento, CA. 71 pp.
  • West, N. E. 1979. Survival patterns of major perennials in salt desert shrub communities of southwestern Utah. Journal of Range Management 32(6):442-445.
  • West, N. E. 1982. Approaches to synecological characterization of wildlands in the Intermountain West. Pages 633-643 in: In-place resource inventories: Principles & practices. A national workshop, University of Maine, Orono. Society of American Foresters, McClean, VA. August 9-14, 1981.
  • West, N. E. 1983b. Intermountain salt desert shrublands. Pages 375-397 in: N. E. West, editor. Temperate deserts and semi-deserts. Ecosystems of the world, Volume 5. Elsevier Publishing Company, Amsterdam.
  • West, N. E., and K. I. Ibrahim. 1968. Soil-vegetation relationships in the shadscale zone of southeastern Utah. Ecology 49(3):445-456.