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G030 Juniperus virginiana - Liriodendron tulipifera / Lonicera tatarica Ruderal Forest Group

Type Concept Sentence: This native ruderal forest group is found on former agricultural or forest plantation sites, or arises from degraded native forest sites, in the cool temperate regions of the eastern United States and Canada.


Common (Translated Scientific) Name: Eastern Red-cedar - Tuliptree / Tatarian Honeysuckle Ruderal Forest Group

Colloquial Name: Eastern North American Native Ruderal Forest

Hierarchy Level:  Group

Type Concept: This native ruderal forest group is found in the cool temperate regions of eastern United States and Canada, from the East Coast, west to the Great Lakes and Tallgrass Prairie region and south to the south-central United States. The vegetation of this group shows evidence of former and heavy human disturbance, particularly to the soils, such as through plowing, grading, skidding, etc., but has otherwise been allowed to succeed more-or-less spontaneously. The vegetation is a somewhat disparate mix of weedy and native species, and the ecological and floristic organization of the vegetation is not clear. The tree layer is dominated (>50% cover) by "weedy" or generalist native tree species, sometimes accompanied by exotic/invasive tree species. The list of ruderal tree species is as follows: Conifers: Juniperus virginiana, Pinus rigida, Pinus strobus, Pinus virginiana. Hardwoods: Acer negundo, Acer rubrum, Amelanchier spp., Betula populifolia, Crataegus spp., Crataegus crus-galli, Crataegus mollis, Diospyros virginiana, Fraxinus americana, Gleditsia triacanthos, Gymnocladus dioicus, Juglans nigra, Liquidambar styraciflua, Liriodendron tulipifera, Morus spp., Malus fusca, Morus rubra, Populus tremuloides, Prunus pensylvanica, Prunus serotina, Prunus americana, and Robinia pseudoacacia. Naturalized exotics include the conifer Pinus thunbergii and hardwoods Acer platanoides and Ailanthus altissima. Robinia pseudoacacia, although a native in the central hardwoods region, is so widely planted outside of its range that it is essentially exotic in character. The weedy natives often form mono-dominant or mixed dominance stands. Understory shrub and herb species vary from exotic invasives to native generalists. Invasive shrub species include a variety of honeysuckles (Lonicera japonica, Lonicera morrowii, Lonicera tatarica, Lonicera x bella), Rhamnus cathartica, and others. Sites show evidence of former and heavy human use, particularly with extensive and intensive soil disturbances, including plowing, grading, skidding, etc. Sites are often on dry-mesic to wet-mesic sites, suitable for agriculture or forest plantations.

Diagnostic Characteristics: A specified list of "weedy" or generalist native species [see Floristics] form mono-dominant stands and typically have associated shrub and herb layers that contain generalist native or exotic species. The weedy natives are >80% (>90%?) of the canopy.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This group is defined by a specified list of generalist native species, sometimes accompanied by naturalized exotics. The concept of this group is being tested by a USGS Forest Inventory and Analysis (FIA) project with NatureServe staff and USFS staff (Faber-Langendoen and Menard 2006). Species that typify the ruderal category include those that are able to establish on disturbed sites, especially when the disturbance includes soil alternation, such as plowing, landfills, graded sites, etc. Species traits include shade intolerance, wind dispersal, and high reproductive capacity. More information on the characteristic dominant trees on these ruderal sites is needed to distinguish this group from ~Southeastern Native Ruderal Forest Group (G031)$$. Species more typical of that group include Carya illinoinensis, Catalpa spp. (including Catalpa bignonioides, Catalpa speciosa), or Maclura pomifera. Where the ground layer is native and the ruderal tree species is native (e.g., Cornus florida), the stand may better fit into a successional phase of a native type. This group could be split into conifer-dominated versus hardwood-dominated stands, or by dry versus mesic sites, or by native versus exotics species. Decisions will be deferred until alliances are better developed.

Native forest plantation stands (tracked in 7. ~Agricultural & Developed Vegetation Cultural Class (CCL01)$$) could become this ruderal type if not intensively managed, as the planted trees begin to die out and the ground layer is invaded by native ruderal species.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Tree canopy varies from hardwood-dominated to conifer-dominated, with open to closed canopy cover. Stands have an irregular structure, though remnants of abandoned forest plantation structure, such as row plantings, may be evident in some cases.

Floristics: The vegetation is a somewhat disparate mix of weedy and native species, and the ecological and floristic organization of the vegetation is not clear. The tree layer is dominated by >80% (>90%?) cover of "weedy" or generalist native tree species, as compared to exotic/invasive tree species. The list of ruderal tree species is as follows: Conifers: Juniperus ashei, Juniperus virginiana, Pinus rigida, Pinus strobus, Pinus virginiana. Hardwoods: Acer negundo, Acer platanoides, Acer rubrum, Ailanthus altissima, Amelanchier spp., Betula populifolia, Crataegus spp., Crataegus crus-galli, Crataegus mollis, Diospyros virginiana, Fraxinus americana, Gleditsia triacanthos, Gymnocladus dioicus, Juglans nigra, Liquidambar styraciflua, Liriodendron tulipifera, Morus spp., Malus fusca, Morus rubra, Populus tremuloides, Prunus pensylvanica, Prunus serotina, Prunus americana, and Robinia pseudoacacia. These species form mono-dominant or mixed dominance stands. Understory shrub and herb species vary from exotic invasives to native generalists. Invasive shrub species include a variety of honeysuckles (Lonicera japonica, Lonicera morrowii, Lonicera tatarica, Lonicera x bella), Rhamnus cathartica, and others.

Dynamics:  Stands most typically represent the tree-dominated stage in the development of vegetation on sites that were heavily disturbed by humans, including plowing, grading, skidding, etc. In these cases, earlier stages of vegetation include annual and perennial weeds, grasslands and shrublands [see ~Eastern North American Ruderal Meadow & Shrubland Group (G059)$$]. Because the extensive soil disturbances typically lead to a mix of weedy native and exotic shrub and herb species, the stands take on a ruderal composition, even if native trees invade or are planted. Canopy cover may be as low as 10%, but eventually stands may have more-or-less continuous canopy, leading to a shift to a more shade-tolerant ground layer.

The successional stages of this type have been described in many studies, particularly the early stages of tree invasion into old fields (e.g., Singleton et al. 2001), as have the limited recovery of these ruderal forests to a composition resembling the historic native forests of the region (Bellemare et al. 2002). See also Wright and Fridley (2010) for the biogeographic variation among stands of this type.

This type may also form in other ways. First, native forest plantation stands (tracked in 7. ~Agricultural & Developed Vegetation Cultural Class (CCL01)$$)) could become ruderal stands if not intensively managed, as the planted trees begin to die out, and the ground layer is invaded by native species. Second, native forest stands that have not been plowed or planted may be stressed to the point where the characteristic native combination of species is altered (Curtis 1959). These stands are probably best tracked as altered variants of native types until the overstory itself is substantially altered.

Environmental Description:  Sites show evidence of former and heavy human use, particularly with extensive and intensive soil disturbances, including plowing, grading, skidding, etc. Sites are often on dry-mesic to wet-mesic sites, suitable for agriculture or forest plantations.

Geographic Range: This native ruderal forest group is found in the cool temperate regions of eastern United States and Canada, from the East Coast, west to the Great Lakes and Tallgrass Prairie region and south to the south-central United States.

Nations: CA,US

States/Provinces:  AL, AR, CT, DC, DE, GA, IA, IL, IN, KS, KY, LA, MA, MB?, MD, ME, MI, MN, MO, MS, NB, NC, ND, NE, NF, NH, NJ, NS, NY, OH, OK, ON, PA, PE, QC, RI, SC, SD, TN, TX, VA, VT, WI, WV




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: GNA

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: No Data Available

Concept Author(s): D. Faber-Langendoen and S. Menard (2006)

Author of Description: D. Faber-Langendoen

Acknowledgements: S. Franklin, J. Vanderhorst

Version Date: 05-05-15

  • Bellemare, J., G. Motzkin, and D. R. Foster. 2002. Legacies of the agricultural past in the forested present: An assessment of historical land-use effects on rich mesic forests. Journal of Biogeography 29:1401-1420.
  • Curtis, J. T. 1959. The vegetation of Wisconsin: An ordination of plant communities. Reprinted in 1987. University of Wisconsin Press, Madison. 657 pp.
  • Faber-Langendoen, D., J. Drake, S. Gawler, M. Hall, C. Josse, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, L. Sneddon, K. Schulz, J. Teague, M. Russo, K. Snow, and P. Comer, editors. 2010-2019a. Divisions, Macrogroups and Groups for the Revised U.S. National Vegetation Classification. NatureServe, Arlington, VA. plus appendices. [in preparation]
  • Faber-Langendoen, D., and S. Menard. 2006. A key to eastern forests of the United States: Macrogroups, groups, and alliances. September 15, 2006. NatureServe, Arlington, VA.
  • Singleton, R., S. Gardescu, P. L. Marks, and M. A. Geber. 2001. Forest herb colonization of postagricultural forests in central New York State, USA. Journal of Ecology 89:325-338.
  • Wright, J. P., and J. D. Fridley. 2010. Biogeographic synthesis of secondary succession rates in eastern North America. Journal of Biogeography 37:1584-1596.