Print Report
G208 Pseudotsuga menziesii - Notholithocarpus densiflorus - Arbutus menziesii Forest Group
Type Concept Sentence: This forested group is characterized by mixes of coniferous and broad-leaved evergreen trees, such as Arbutus menziesii, Chrysolepis chrysophylla, Notholithocarpus densiflorus, Pseudotsuga menziesii, Quercus chrysolepis, and Umbellularia californica, found in coastal California mountains.
Common (Translated Scientific) Name: Douglas-fir - Tanoak - Pacific Madrone Forest Group
Colloquial Name: Californian Moist Coastal Mixed Evergreen Forest
Hierarchy Level: Group
Type Concept: This forest group is characterized by mixes of coniferous and broad-leaved evergreen trees. Characteristic trees include Arbutus menziesii, Chrysolepis chrysophylla, Notholithocarpus densiflorus, Pseudotsuga menziesii, Quercus chrysolepis, and Umbellularia californica. Pinus coulteri occurs in scattered stands. On the eastern fringe of this group, in the western Klamath Mountains, other conifers occur such as Calocedrus decurrens, Pinus ponderosa, and Chamaecyparis lawsoniana. In the southern portion of the range, stands are a mixture of Abies bracteata and Quercus chrysolepis. More northerly stands tend to have dense or diverse shrub understories, with Corylus cornuta, Gaultheria shallon, Mahonia nervosa, Quercus sadleriana, Rhododendron macrophyllum, Toxicodendron diversilobum, and Vaccinium ovatum being common. This group occurs in California in the Santa Cruz Mountains, the Santa Lucia Mountains, throughout the outer and middle Coast Ranges, south across the outer central Coast Ranges and north in the northern Coast Ranges and western Klamath Mountains, into southwestern Oregon, and locally in the northern Sierra Nevada. It can occur on metasediments and granitic substrate, on Franciscan Formation soils (metasedimentary sandstones, schists and shales) with moderate to high rainfall in the Coast Ranges. Historic fire frequency in this group was higher than for redwood-dominated systems (every 50-100 years).Southern stands have a sparser shrub understory; Toxicodendron diversilobum is the most constant shrub, along with much Polystichum munitum. Especially in the south, stands are restricted to fire-protected sites (extremely steep, northerly, mesic slopes and coves) where fires from adjacent chaparral systems do not carry. Historic fire frequency was likely higher than the north. Stands with just conifers present are a part of ~Californian Conifer Forest & Woodland Group (G198)$$.
Diagnostic Characteristics: This forested group is characterized by mixes of coniferous and broad-leaved evergreen trees. Characteristic trees include Pseudotsuga menziesii, Pseudotsuga macrocarpa, Quercus chrysolepis, Notholithocarpus densiflorus, Arbutus menziesii, Umbellularia californica, and Chrysolepis chrysophylla var. chrysophylla.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: In northern California, especially around Point Reyes, there are stands dominated by Umbellularia californica. These nearly pure stands are a part of this group, as it is a disturbance-driven species and grows rapidly with full sunlight. With time and succession, other trees will succeed into the canopy. This is in contrast to small patches or individuals of Umbellularia californica in some of the various chaparral groups. Here there are no chaparral shrubs in the understory. Notholithocarpus densiflorus in mixed stands of various pines and oaks is the indicator species for this group in many places in the Coast Ranges throughout northern and central California, and locally in the northern Sierra Nevada.
It has been proposed by the reviewer to limit this group to the distribution of Notholithocarpus densiflorus and Arbutus menziesii, with Pseudotsuga menziesii. The southerly stands with Pseudotsuga macrocarpa (and lacking Notholithocarpus densiflorus or Arbutus menziesii) do not belong in this group. It is suggested to have a more southerly "mixed evergreen" group. For now this represents a more limited group concept. Pseudotsuga macrocarpa stands have been moved to ~Californian Montane Conifer Forest & Woodland Group (G344)$$, which is where Sawyer et al. (2009) place them, along with the Abies bracteata stands.
It has been proposed by the reviewer to limit this group to the distribution of Notholithocarpus densiflorus and Arbutus menziesii, with Pseudotsuga menziesii. The southerly stands with Pseudotsuga macrocarpa (and lacking Notholithocarpus densiflorus or Arbutus menziesii) do not belong in this group. It is suggested to have a more southerly "mixed evergreen" group. For now this represents a more limited group concept. Pseudotsuga macrocarpa stands have been moved to ~Californian Montane Conifer Forest & Woodland Group (G344)$$, which is where Sawyer et al. (2009) place them, along with the Abies bracteata stands.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: Forests generally have closed or near-closed canopies, often in older stands with two strata in the canopy, 15-35 m tall, and often with a dense shrub understory structure consisting of many regenerating canopy tree species.
Floristics: It is characterized by mixes of coniferous and broad-leaved evergreen trees. Characteristic trees include Arbutus menziesii, Chrysolepis chrysophylla var. chrysophylla, Notholithocarpus densiflorus (= Lithocarpus densiflorus), Pseudotsuga menziesii, Quercus chrysolepis, Umbellularia californica. Pinus coulteri occurs in scattered stands. On the eastern fringe of this group, in the western Klamath Mountains, other conifers occur such as Calocedrus decurrens, Pinus ponderosa, and Chamaecyparis lawsoniana. In the southern portion of the range, stands are a mixture of Abies bracteata and Quercus chrysolepis, and Notholithocarpus densiflorus, Arbutus menziesii, Umbellularia californica, and Chrysolepis chrysophylla become less important. More northerly stands tend to have dense or diverse shrub understories, with Corylus cornuta, Gaultheria shallon, Mahonia nervosa, Quercus sadleriana, Rhododendron macrophyllum, Toxicodendron diversilobum, and Vaccinium ovatum being common. Southern stands have a sparser shrub understory; Toxicodendron diversilobum is the most constant shrub, with Ribes spp. occasionally present, along with much Polystichum munitum.
Dynamics: Especially in the south, stands are restricted to fire-protected sites (extremely steep, northerly, mesic slopes and coves) where fires from adjacent chaparral systems do not carry. Historic fire frequency was likely higher than the north.
Environmental Description: This group occurs just inland from the redwood belt of this region. It can occur on metasediments and granitic substrate, on Franciscan Formation soils (metasedimentary sandstones, schists and shales) in areas with moderate to high rainfall on the Coast Ranges of California. Especially in the south, stands are restricted to fire-protected sites (extremely steep, northerly, mesic slopes and coves) where fires from adjacent chaparral systems do not carry.
Geographic Range: This group occurs in California from the Santa Lucia and Santa Cruz mountains north to the North Coast Ranges and western Klamath Mountains into southwestern Oregon, and locally in the Sierra Nevada. It also occurs in localized areas of the central to northern Sierra Nevada and southern and eastern Klamath Mountains.
Nations: US
States/Provinces: CA, OR
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.833214
Confidence Level: High
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.1 Warm Temperate Forest & Woodland Formation | F018 | 1.B.1 |
Division | 1.B.1.Nc Californian Forest & Woodland Division | D007 | 1.B.1.Nc |
Macrogroup | 1.B.1.Nc.1 California Live Oak - California Laurel - Western Cypress species Forest & Woodland Macrogroup | M009 | 1.B.1.Nc.1 |
Group | 1.B.1.Nc.1.c Douglas-fir - Tanoak - Pacific Madrone Forest Group | G208 | 1.B.1.Nc.1.c |
Alliance | A0106 Douglas-fir - Tanoak Forest Alliance | A0106 | 1.B.1.Nc.1.c |
Alliance | A3357 Tanoak - Pacific Madrone Forest Alliance | A3357 | 1.B.1.Nc.1.c |
Concept Lineage: G199 merged into G208
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: > Arbutus menziesii (Madrone forest) Alliance (Sawyer et al. 2009) [73.200.00]
> Lithocarpus densiflorus (Tanoak forest) Alliance (Sawyer et al. 2009) [73.100.00]
> Pseudotsuga menziesii - Lithocarpus densiflorus (Douglas fir - tanoak forest) Alliance (Sawyer et al. 2009) [82.500.00]
= Douglas-fir - Tanoak - Pacific Madrone: 234 (Eyre 1980)
> Lithocarpus densiflorus (Tanoak forest) Alliance (Sawyer et al. 2009) [73.100.00]
> Pseudotsuga menziesii - Lithocarpus densiflorus (Douglas fir - tanoak forest) Alliance (Sawyer et al. 2009) [82.500.00]
= Douglas-fir - Tanoak - Pacific Madrone: 234 (Eyre 1980)
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