Print Report

G009 Pinus palustris / Quercus margarettae / Aristida spp. Woodland Group

Type Concept Sentence: This group represents stands of Pinus palustris with scattered tall shrubs or small trees of Quercus marilandica, Quercus incana, Quercus margarettae, or Quercus falcata, and a grass-dominated understory often with Schizachyrium scoparium, Aristida stricta, or Aristida beyrichiana on sandy to loamy soils of uplands in the southern coastal plain.


Common (Translated Scientific) Name: Longleaf Pine / Sand Post Oak / Three-awn species Woodland Group

Colloquial Name: Dry-Mesic Loamy Longleaf Pine Woodland

Hierarchy Level:  Group

Type Concept: This group represents stands of Pinus palustris on sandy to loamy soils on uplands ranging from gently rolling, broad ridgetops to steeper sideslopes, and in mesic swales and terraces. The vegetation is generally open woodlands, with irregularly scattered trees of Pinus palustris, and usually clumps of midstory Quercus spp. and a grassy understory. There tends to be a fairly high diversity of forbs (broadleaf herbaceous plants), especially in sites that have been burned frequently (i.e., three or more times per decade). This group does not include the xeric and subxeric Pinus palustris - Quercus laevis habitats. It is found from southeastern Virginia to east Texas, including most of Florida. It does not occur in the Mississippi Alluvial Plain.

Diagnostic Characteristics: This group represents stands of Pinus palustris on loamy sand or sandy loam soils on upland sites. This group does not include the dry Pinus palustris - Quercus laevis habitats, but represents those that have moderate moisture and nutrient availability.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Classification units below the group such as NVC alliances representing vegetation dominated by Pinus palustris can be subdivided by biogeography, from northeast to southwest across the coastal plains from Virginia to Texas. Longleaf pine-dominated stands in the rocky submontane areas of the Piedmont as well as the Ridge and Valley (from North Carolina to Alabama) are classified as a separate forest group. This group includes upland, not wet, longleaf pine habitats.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: These are open woodlands; the trees are primarily needle-leaved evergreen conifers and on loamy soils can be straight, well-formed, and moderately tall. Open canopies and grass-dominated understories are typical of sites managed with prescribed fire. Sites lacking a hardwood midstory were more prevalent prior to the twentieth century.

Floristics: This vegetation is naturally dominated by Pinus palustris. Scrub oaks, such as Quercus marilandica, Quercus incana, Quercus margarettae, Quercus falcata, and sometimes Quercus laevis form a sparse or clumped understory in most stands, in fact in all but the most mesic ones. Low shrubs, mostly ericaceous, may be abundant. East of the Mississippi River, Aristida stricta (in North and South Carolina) or Aristida beyrichiana (from South Carolina to Mississippi) are usually the dominant or at least a characteristic herb. In central South Carolina and west of the Mississippi River, both Aristida beyrichiana and Aristida stricta are absent and various other grasses dominate. Many examples have moderate to high plant species richness, with most of the species in the herb layer. Some mesic associations have high species richness values measured at the 1/10-hectare scale. Forbs, especially composites and legumes, are usually also an important herb component. Some typical mesic to dry-mesic grass species include Andropogon gyrans var. gyrans, Andropogon ternarius, Andropogon virginicus, Panicum virgatum, Schizachyrium scoparium, Schizachyrium tenerum, Sorghastrum elliottii, Sorghastrum nutans, Sorghastrum secundum, Sporobolus clandestinus, and Sporobolus junceus.

Dynamics:  Frequent fire is the predominant natural disturbance of this group, and it is dependent on frequent prescribed fire every 2-3 years. Fires are mostly low to moderate in intensity, removing above-ground vegetation of herbs and shrubs, but have little effect on the fire-tolerant Pinus palustris trees. Vegetation recovers very quickly from fire, with live herbaceous biomass often restored in just a few weeks after a growing season fire. Many plants have their flowering triggered by burning. In the absence of fire, species which are less able to withstand fire increase. Quercus spp. and other shrubs, kept to low density and mostly reduced to shrub size by fire, become tall and dense and can reduce Pinus palustris tree regeneration. Herb layer density and diversity declines in the absence of fire. On most soils, vegetation undergoes substantial structural alteration and reduction in species richness with lack of fire. Canopies are multi-aged, consisting of a fine mosaic of small even-aged patches driven by gap-phase regeneration. Pinus palustris is shade-intolerant and slow to reach reproductive age but is very long-lived, and continues to produce more cones as it grows older, after age 100. Some insect populations recolonize burned areas from nearby unburned patches. Sites managed with late spring or growing season prescribed fire provide high arthropod biomass (James et al. 2001, Taylor 2003).

Environmental Description:  The vegetation of this group occurs on well-drained sandy to loamy soils on upland sites of the coastal plain, on landforms that include loamy to sandy flats, relict beach system deposits, eolian sand deposits, Carolina bay rims (Bennett and Nelson 1991), and low rolling hills. Climate: Longleaf pine grows in humid, warm temperate climates characterized by hot summers and mild winters. Annual mean temperatures range from 16-23°C (60-74°F). Annual precipitation ranges from 109 to 175 cm (43-69 inches) (Boyer 1990). Fall is the driest season of the year, although periods of drought during the growing season are not unusual (Boyer 1990). Soil/substrate/hydrology: Soils range from mesic to dry and from sandy to loamy or occasionally clayey. Most natural remnants are on sand, but many examples probably once occurred on loamy soils. Soils are largely acidic and infertile, though local richer, mesic sites occur. Ultisols most commonly associated with longleaf pine are the Typic Paleudults and Plinthic Paleudults. Some areas are occupied by Psamments and other coarser-textured soils.

Geographic Range: The vegetation of this group is found from southeastern Virginia to east Texas, including most of Florida. It does not occur in the Mississippi Alluvial Plain.

Nations: US

States/Provinces:  AL, FL, GA, LA, MS, NC, SC, TX, VA




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: > Dry Upland Longleaf Pine Woodlands (Edwards et al. 2013)
? Grossarenic Dry Uplands (Turner et al. 1999)
> Longleaf-Bluestem Uplands (Ajilvsgi 1979)
> Mesic Upland Longleaf Pine Woodlands (Edwards et al. 2013)
> Sandhill Pine Forest (Marks and Harcombe 1981)
= Upland Pine (FNAI 2010a)
? Upland Pine Forest (Marks and Harcombe 1981)
> Xeric Sandhill Scrub (Bennett and Nelson 1991)
> Xeric stream terrace sand ridge subtype (of Upland Longleaf Pine Savanna) (Bridges and Orzell 1989a)

Concept Author(s): W.G. Wahlenberg (1946)

Author of Description: C.W. Nordman

Acknowledgements: No Data Available

Version Date: 05-12-15

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