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CEGL004143 Lasallia papulosa - Stereocaulon glaucescens - Chrysothrix chlorina Nonvascular Vegetation

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Common Toadskin Lichen - Snow Lichen - Sulphur Dust Lichen Nonvascular Vegetation

Colloquial Name: Central Appalachian Mafic Boulderfield

Hierarchy Level:  Association

Type Concept: This association is known only from four counties on the northern Virginia Blue Ridge, where it occurs on fully exposed, minimally weathered metabasalt boulderfields at elevations from about 670 to 1160 m (2200-3800 feet). Vascular plants are generally absent and lichens dominate these habitats. Lasallia papulosa and Stereocaulon glaucescens are generally abundant and conspicuous in variable combinations. Although not abundant, Chrysothrix chlorina is scattered on sheltered boulder faces and in grottoes and is a good diagnostic species as it appears to be absent from siliciclastic boulderfields in this region. A variety of other foliose, crustose and fruticose lichen species are associated, including several characteristic arctic-boreal species at higher elevations.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Classification of this association is based on geographically limited lichen inventories in Shenandoah National Park. In the park inventory, lichens were mass-collected from boulderfield and outcrop habitats on different geologic substrates and specimens were identified by Richard Harris (New York Botanical Garden), Don Flenniken (author of Macrolichens of West Virginia), and James Lawry (George Mason University). Classification of lichen communities in eastern North America is currently difficult and tentative because inventory and data are generally lacking. Because Stereocaulon glaucescens and many other species in this association have northern or arctic-boreal ranges, related boulderfield communities are most likely to be found north of Virginia.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: Vascular plants are generally absent and lichens dominate. Maximum patch size is about one acre, and many patches are much smaller. Lasallia papulosa and Stereocaulon glaucescens are generally abundant and conspicuous in variable combinations. Although not abundant, Chrysothrix chlorina is a good diagnostic species as it appears to be absent from siliciclastic and granitic boulderfields in this region; it is scattered on sheltered boulder faces and in grottoes, often with Psilolechia lucida, Usnea halei, and Ramalina intermedia. Other minor umbilicate and foliose species include Flavoparmelia baltimorensis, Parmelia sulcata, and Umbilicaria muhlenbergii. Many crustose species occur, including Aspicilia cinerea, Diploschistes scruposus, Fuscidea recensa, Lepraria spp., Porpidia spp., Rhizocarpon rubescens, and Trapeliopsis granulosa. Flat surfaces and interstices that have thin deposits of organic matter often support a variety of fruticose lichens, including Cladonia rangiferina (= Cladina rangiferina), Cladonia stellaris (= Cladina stellaris), Cladonia crispata, Cladonia furcata, Cladonia pleurota, and Cladonia squamosa. At higher elevations, a number of characteristic arctic-boreal lichens occur, including Cladonia coccifera, Melanelia stygia, Microcalicium arenarium, Parmelia omphalodes, Porpidia tuberculosa, Rhizocarpon geographicum, and Lasallia caroliniana (= Umbilicaria caroliniana). Along the edges of the boulderfields, scattered individuals or patches of Polypodium appalachianum, Hylotelephium telephioides, and other vascular plants may occur in transition zones with forests or woodlands.

Dynamics:  The exposed rock surfaces supporting this community are subject to fluctuating, daily extremes of temperature, humidity, and moisture saturation during the growing season, as well as to low temperatures, high winds, and frequent ice during the winter. Boulderfield habitats have resulted from periglacial phenomena and the collapse of resistant strata from weathering and erosion of weaker underlying rocks. Most Central Appalachian boulderfields are well-weathered and now support woodland or forest vegetation. In this region, metabasalt underlies much less acreage than quartzite and sandstone and is less resistant to weathering. Consequently, exposed metabasalt boulderfields are quite rare.

Environmental Description:  This association occurs on fully exposed, minimally weathered metabasalt boulderfields at middle and high elevations of the northern Blue Ridge. The known elevation range is from about 670 to 1160 m (2200-3800 feet), with the majority of acreage located above 915 m (3000 feet). Aspect is variable among sites, but slopes are typically steep to very steep (often >30°). Block size is typically <1 m, with the surficial boulders in the field somewhat loose. Although this association is most extensive on boulder deposits, it may also occur on fully exposed outcrops associated with the boulderfields.

Geographic Range: This association is known only from metabasalt districts in four counties on the northern Virginia Blue Ridge. The potential range is larger, but suitable habitat (i.e., mafic substrates at higher elevations) are not known to occur elsewhere in the Central Appalachians.

Nations: US

States/Provinces:  VA




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: G1?

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Lasallia papulosa - Stereocaulon glaucescens - Chrysothrix chlorina Nonvascular Vegetation (Fleming et al. 2007a)

Concept Author(s): G.P. Fleming et al. (2007a)

Author of Description: G.P. Fleming

Acknowledgements: No Data Available

Version Date: 12-07-06

  • Fleming, G. P., A. Belden, Jr., K. E. Heffernan, A. C. Chazal, N. E. Van Alstine, and E. M. Butler. 2007a. A natural heritage inventory of the rock outcrops of Shenandoah National Park. Unpublished report submitted to the National Park Service. Natural Heritage Technical Report 07-01. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 433 pp. plus appendixes.
  • Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
  • Flenniken, D. G. 1999. The macrolichens in West Virginia. Carlisle Printing, Walnut Creek. 231 pp. plus 26 color plates.
  • Southeastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Durham, NC.
  • Young, J., G. Fleming, W. Cass, and C. Lea. 2009. Vegetation of Shenandoah National Park in relation to environmental gradients, Version 2.0. Technical Report NPS/NER/NRTR--2009/142. National Park Service, Philadelphia, PA. 389 pp.