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CEGL004142 Lasallia (papulosa, pensylvanica) - Dimelaena oreina - (Melanelia culbersonii) Nonvascular Vegetation

Type Concept Sentence: This acidic boulderfield natural community of the Central Appalachians consists of lichen-covered boulders on steep slopes; the foliose lichens Lasallia papulosa and Lasallia pensylvanica are usually abundant.


Common (Translated Scientific) Name: (Common Toadskin Lichen, Pennsylvania Toadskin Lichen) - Golden Moonglow Lichen - Culberson''s Black-parmelia Nonvascular Vegetation

Colloquial Name: Central Appalachian Acidic Boulderfield

Hierarchy Level:  Association

Type Concept: This association is widely but locally distributed from the western Piedmont foothills in Maryland and Virginia through the Blue Ridge and Ridge and Valley portions of the Central Appalachians, north at least to the Delaware Water Gap in Pennsylvania and New Jersey. It occurs primarily on fully exposed, minimally weathered quartzite and sandstone boulderfields at elevations from about 300 to 1000 m (1000-3300 feet). On the largest occurrences, vascular plants are generally absent and lichens dominate these habitats. Lasallia papulosa and Lasallia pensylvanica, either singly or in combination, are generally abundant and conspicuous. Dimelaena oreina abundantly covers many dry, exposed rock surfaces that are not covered with Lasallia spp. and larger foliose lichens. Although of scattered occurrence, Melanelia culbersonii has been found across the full elevation range of the type and is a good diagnostic species, as it appears to be restricted mostly to siliciclastic rocks (and occasionally coarse-grained, quartz-rich granites that are nearly devoid of dark minerals) in this region. A variety of other foliose, crustose and fruticose lichen species are associated. Smaller, more marginal occurrences have sparse vascular plant cover, primarily stunted trees of Betula lenta, Sassafras albidum, and Quercus montana, ericaceous shrubs, and scrambling vines of Parthenocissus quinquefolia.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Classification of this association is based on geographically limited lichen inventories in Shenandoah National Park, but it is believed to be widely applicable to similar boulderfields that are characteristic of Central Appalachian siliciclastic ridges. In the Shenandoah National Park inventory, lichens were mass-collected from boulderfield and outcrop habitats on different geologic substrates, and specimens were identified by Richard Harris (New York Botanical Garden), Don Flenniken (author of Macrolichens of West Virginia), and James Lawry (George Mason University). Classification of lichen communities in eastern North America is currently difficult and tentative because inventory and data are generally lacking. The classification of this type versus ~Betula lenta - Quercus montana / Parthenocissus quinquefolia Woodland (CEGL006565)$$ can be tricky where there is a continuous gradation of vegetation cover (as at Delaware Water Gap), but generally this is applied where there is less than 25% vascular vegetation and nonvascular species are dominant.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: On the largest occurrences, vascular plants are generally absent and lichens dominate. Maximum patch size is about ten acres, but most patches are considerably smaller. Lasallia papulosa and Lasallia pensylvanica, either singly or in combination, are generally abundant and conspicuous. Dimelaena oreina abundantly covers many dry, exposed rock surfaces that are not covered with Lasallia spp. and larger foliose lichens. Although of scattered occurrence, Melanelia culbersonii has been found across the full elevation range of the type and is a good diagnostic species, as it appears to be restricted mostly to siliciclastic rocks (and occasionally coarse-grained, quartz-rich granites that are nearly devoid of dark minerals) in this region. Other minor umbilicate and foliose species include Hypogymnia physodes, Physcia subtilis, Umbilicaria muhlenbergii, Xanthoparmelia conspersa, and Xanthoparmelia plittii. Many crustose species occur, including Aspicilia cinerea, Fuscidea recensa, Lecanora spp., Lepraria spp., Rhizocarpon reductum, and Sarcogyne clavus. Flat surfaces and interstices that have thin deposits of organic matter often support a variety of fruticose lichens, including Cladonia rangiferina (= Cladina rangiferina), Cladonia uncialis (= Cladina uncialis), Cladonia crispata, Cladonia macilenta, Cladonia ochrochlora, and Cladonia squamosa. In the upper elevation range, boreal lichens such as Melanelia stygia and Arctoparmelia centrifuga are present, but they are not abundant. Along the edges of the boulderfields, scattered individuals of Parthenocissus quinquefolia, Vaccinium spp., and other vascular plants may occur in transition zones with forests or woodlands. Smaller, more marginal occurrences frequently have sparse vascular plant cover, primarily stunted trees of Betula lenta, Sassafras albidum, Quercus montana (= Quercus prinus), Quercus coccinea, Quercus velutina, Carya glabra, and Carya ovalis. Widely scattered shrubs may include Kalmia latifolia and other ericads. Herbs are usually absent, but Dicentra eximia is known from some occurrences.

Dynamics:  The exposed rock surfaces supporting this community are subject to fluctuating, daily extremes of temperature, humidity, and moisture saturation during the growing season, as well as to low temperatures, high winds, and frequent ice during the winter. Boulderfield habitats have resulted from periglacial phenomena and the collapse of resistant strata from weathering and erosion of weaker underlying rocks. Most Central Appalachian boulderfields are well-weathered and now support woodland or forest vegetation. Relatively few remain fully exposed and support wholly nonvascular communities.

Environmental Description:  This association occurs primarily on fully exposed, minimally weathered quartzite and sandstone boulderfields at low and middle elevations of the Northern Blue Ridge, Ridge and Valley, and adjacent foothills of the upper Piedmont. A few occurrences have also been noted on boulderfields composed of acidic granitic rocks (e.g., on Old Rag Mountain in Shenandoah National Park). The known elevation range is from about 300 to 1000 m (1000-3300 feet). Aspect is variable among sites, but slopes are typically steep to very steep (often >30°). Block size is highly variable, from relatively small and loose stones (<1 m in diameter) to large, stable boulders (>1 m in diameter). Although this association is most extensive on boulder deposits, it may also occur on outcrops associated with the boulderfields, or on very large, exposed cliffs. There is little or no available soil except for occasional small deposits of organic matter in crevices.

Geographic Range: This association is widely but locally distributed from the western Piedmont foothills in Maryland (e.g., Sugarloaf Mountain) and Virginia (e.g., Bull Run Mountain) through the Blue Ridge and Ridge and Valley portions of the Central Appalachians, north at least to the Delaware Water Gap in Pennsylvania and New Jersey. The type is fairly common on the siliciclastic western flank of the Northern Blue Ridge and throughout the Ridge and Valley in west-central and northwestern Virginia. It is also frequent in the Ridge and Valley region of Pennsylvania (T. Smith pers. comm.) but may be restricted to the northeastern tier of counties in West Virginia. The potential range of this association covers a much larger geographic area.

Nations: US

States/Provinces:  MD, NJ, PA, VA, WV




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: G5

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Lasallia (papulosa, pensylvanica) - Dimelaena oreina - (Melanelia culbersonii) Nonvascular Vegetation (Fleming et al. 2007a)

Concept Author(s): G.P. Fleming et al. (2007a)

Author of Description: G.P. Fleming

Acknowledgements: No Data Available

Version Date: 12-07-06

  • Eastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boston, MA.
  • Fleming, G. P., A. Belden, Jr., K. E. Heffernan, A. C. Chazal, N. E. Van Alstine, and E. M. Butler. 2007a. A natural heritage inventory of the rock outcrops of Shenandoah National Park. Unpublished report submitted to the National Park Service. Natural Heritage Technical Report 07-01. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 433 pp. plus appendixes.
  • Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
  • Flenniken, D. G. 1999. The macrolichens in West Virginia. Carlisle Printing, Walnut Creek. 231 pp. plus 26 color plates.
  • Perles, S. J., G. S. Podniesinski, E. Eastman, L. A. Sneddon, and S. C. Gawler. 2007. Classification and mapping of vegetation and fire fuel models at Delaware Water Gap National Recreation Area. Technical Report NPS/NER/NRTR--2007/076. National Park Service, Philadelphia, PA. 2 volumes.
  • Smith, T. Personal communication. Coordinator. Virginia Natural Heritage Program, Richmond VA.
  • WVNHP [West Virginia Natural Heritage Program]. No date. Unpublished data. West Virginia Natural Heritage Program, Elkins.
  • Young, J., G. Fleming, W. Cass, and C. Lea. 2009. Vegetation of Shenandoah National Park in relation to environmental gradients, Version 2.0. Technical Report NPS/NER/NRTR--2009/142. National Park Service, Philadelphia, PA. 389 pp.