Print Report

CEGL005912 Abies lasiocarpa - Picea engelmannii / Clintonia uniflora Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Subalpine Fir - Engelmann Spruce / Bride''s Bonnet Forest

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This association is broadly distributed across the inland northwest and northern Rocky Mountains, where it is associated with a maritime component of climate. It occurs as far east as the Continental Divide in northwestern Montana and Alberta, extending only a few air miles to the east of this strongest of topographic breaks. This community occupies relatively moist (mesic) and cool sites having free air drainage and lacking frost pocket conditions. Elevations range from 1066 to 1710 m (3500-5600 feet). It occurs on slopes of all degrees of steepness and aspect orientation, though it is more likely to occur from toeslope through midslope positions (predominantly collecting positions). At the dry extremes of its distribution it is more strongly associated with protected positions such as concave slopes, moist depressions in gently sloping plateaus, stringers along perennial stream bottoms, toeslopes and northeastern aspects. It occurs on a variety of parent materials, as varying as granite, limestone, glacial-fluvial material, and volcanic ash caps. Abies lasiocarpa and Picea engelmannii dominate the tree canopy. The most important seral species are Pseudotsuga menziesii, Larix occidentalis, and Pinus contorta, however, their cover seldom exceeds 10-15%. Total tree canopy cover is generally in excess of 55% ranging to 75%. Abies lasiocarpa dominates the tree regeneration layer, whereas Picea engelmannii reproduction is more sporadic. The shrub layer ranges from low diversity and cover to extremely diverse and high in cover; the shrubs present are judged largely seral. Those of highest constancy include Acer glabrum, Ribes lacustre, Lonicera utahensis, Rubus parviflorus, Symphoricarpos albus, and Vaccinium membranaceum. The low-shrub component is well-represented by Linnaea borealis, Spiraea betulifolia, Paxistima myrsinites, and, in the northern portion of the type''s range, Vaccinium myrtillus. The only graminoids consistently present are Bromus vulgaris and Calamagrostis rubescens; the latter species may be relatively abundant on early seral stands but with an increase in tree canopy cover it is less abundant. The forb component is relatively species-rich and abundant; cover of the diagnostic species Clintonia uniflora or Tiarella trifoliata generally does not exceed 20%. There are many other forbs with high constancy and the potential to have appreciable cover, including Maianthemum stellatum, Arnica cordifolia, Thalictrum occidentale, and Eucephalus engelmannii; forbs merely with high constancy include Orthilia secunda, Galium triflorum, Goodyera oblongifolia, Osmorhiza berteroi, and Viola orbiculata.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This association is essentially a renamed (including Picea engelmannii) subset of the Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977, Cooper et al. 1987) or the similarly named climax plant association (Johnson and Simon 1987, Williams et al. 1995). It is a subset because previous authors have used Abies lasiocarpa as an indicator species (present and reproducing successfully or having some minimum amount of cover, e.g., 10% (Williams et al. 1995)) which creates a very broad category, capable of being dominated by any number of seral tree species. The type concept used here is that Picea engelmannii or Abies lasiocarpa must have a substantial presence (at least 10% canopy cover, one or both species) but need not be dominant. This interpretation then allows for recognition of stands dominated by seral tree species that occur within this moderate portion of the subalpine zone, as distinct plant associations. The undergrowth concept for the type was kept intact, recognizing the broadest interpretation of Cooper et al. (1987) that either Clintonia uniflora or Tiarella trifoliata need be present and not confined to microsites within the stand for type recognition. This approach essentially recognizes the Clintonia uniflora Phase of Abies lasiocarpa / Clintonia uniflora Habitat Type (Pfister et al. 1977, Cooper et al. 1987), but with other phases split off (note one exception below) as associations unto themselves (e.g., ~Abies lasiocarpa - Picea engelmannii / Menziesia ferruginea / Clintonia uniflora Forest (CEGL005893)$$), it would be redundant to append another Clintonia uniflora to the name. The Aralia nudicaulis Phase was combined with the Clintonia uniflora Phase because after examining data from many stands a considerable overlap was found between the two phases and the unique habitat (rich benchlands) once claimed for the Aralia nudicaulis Phase was not substantiated. Furthermore the Aralia nudicaulis Phase was absent from sites that appeared in all respects to be quintessential habitat, adding doubt to its consistency as an indicator. The existence of this type in the vicinity of the Cascade Crest needs exploration.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: A highly variable mix of Abies lasiocarpa and Picea engelmannii dominate the overstory, ostensibly at all stages of succession; tree canopy cover is generally in excess of 55% ranging to 75%. Certainly the regeneration layers are Abies lasiocarpa-dominated whereas Picea engelmannii reproduction is more sporadic, but its autecological characteristics provide for its presence throughout the sere. The most important seral species are Pseudotsuga menziesii, Larix occidentalis, and Pinus contorta, however, their cover seldom exceeds 10-15% [see Global Classification Comments]. The shrub layer ranges from virtually nonexistent to extremely diverse and high in cover; the shrubs present are judged largely seral in their response and not specific enough in indicator value to be useful in further subdividing this syntaxon. Those of highest constancy include Acer glabrum, Ribes lacustre, Lonicera utahensis, Rubus parviflorus, Symphoricarpos albus, and Vaccinium membranaceum. The low-shrub component is well-represented by Linnaea borealis, Spiraea betulifolia, Paxistima myrsinites, and, in the northern portion of the type''s range, Vaccinium myrtillus. The only graminoids consistently present are Bromus vulgaris and Calamagrostis rubescens; the latter species may be relatively abundant on early seral stands but with an increase in tree canopy cover it is less abundant. The forb component is relatively species-rich and abundant; cover of the diagnostic species Clintonia uniflora or Tiarella trifoliata generally does not exceed 20% [see Global Classification Comments]. A host of forbs with high constancy and the potential to have appreciable cover include Maianthemum stellatum, Arnica cordifolia, Thalictrum occidentale, and Eucephalus engelmannii (= Aster engelmannii); forbs merely exhibiting high constancy include Orthilia secunda, Galium triflorum, Goodyera oblongifolia, Osmorhiza berteroi, and Viola orbiculata.

Dynamics:  It has not been established why particular stands may regenerate following disturbance to dominance by the tree species generally regarded as dominant later in the sere and persisting to long-term stability. It is known that clearcutting followed by broadcast burning may create persistent shrubfields, which retard regeneration (Williams et al. 1995); however, it may be that these shrubfields retard the regeneration of seral tree species more effectively than they do shade-tolerant climax species. With the death of canopy tree species and slow ingrowth of Abies lasiocarpa and Picea engelmannii, other associations, whose undergrowth is characterized by Clintonia uniflora (e.g., Pinus contorta / Clintonia uniflora), will succeed to this association; for stands dominated by Larix occidentalis or Pseudotsuga menziesii this replacement will not occur (statistically speaking) within historic fire-return intervals within this subalpine zone (80-150 years).

Environmental Description:  This community occupies relatively moist (mesic) and cool sites having free air drainage and lacking frost pocket conditions. It is speculated to have the most moderate temperature regimes within ~Abies lasiocarpa - Picea engelmannii Rocky Mountain Moist Forest Alliance (A3614)$$. It occurs on slopes of all degrees of steepness and aspect orientation, though it is more likely to occur from toeslope through midslope positions (predominantly collecting positions). At the dry extreme of its distribution, it is more strongly associated with protected positions such as concave slopes, moist depressions in gently sloping plateau areas, stringers along perennial stream bottoms, toeslopes and northeastern aspects. In the north it ranges from 1066 to 1585 m (3500-5200 feet) (extreme outliers at 1710 m [5600 feet]), whereas to the south it ranges from 1555 to 1710 m (5100-5600 feet). A variety of parent materials are represented, including those as disparate as granite and limestone, including all manner of glacio-fluvial material. In northern Idaho and northwestern Montana it is routinely found on ash caps, ranging from 3 to 60 cm in depth. The soil textures are predominantly loams and silt loams (reflecting in part the volcanic ash); soils typically have less than 15% coarse fragment content and are well-drained.

Geographic Range: This association is broadly distributed across the inland northwest and northern Rocky Mountains, associated with a maritime component of climate. It occurs as far east as the Continental Divide in northwestern Montana and Alberta, but extends only a few air miles to the east of this strongest of topographic breaks.

Nations: CA,US

States/Provinces:  AB, ID, MT, OR, WA




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: G5

Greasons: No Data Available


Concept Lineage: This association (CEGL000307) was based on a Habitat Type concept, including the phases of the H.T. in this association. Phases of the H.T. have now been split out into association elements, as well as the seral components of the H.T.

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Abies lasiocarpa - Picea engelmannii / Clintonia uniflora Forest (Hop et al. 2007)
< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Abies lasiocarpa / Cornus canadensis Plant Association (Williams et al. 1995)
>< Abies lasiocarpa / Linnaea borealis Plant Association (Williams et al. 1995)
< Abies lasiocarpa / Pachistima myrsinites Habitat Type (Daubenmire and Daubenmire 1968)
= Abies lasiocarpa / Pachistima myrsinites Habitat Type (Ogilvie 1962)
>< C69: Picea engelmannii - Abies lasiocarpa - (Pseudotsuga menziesii) / Arnica cordifolia Vegetation Type (Achuff et al. 2002)

Concept Author(s): Hop et al. (2007)

Author of Description: S.V. Cooper

Acknowledgements: No Data Available

Version Date: 02-13-04

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