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CEGL005897 Abies lasiocarpa - Picea engelmannii / Menziesia ferruginea / Streptopus amplexifolius Woodland

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Subalpine Fir - Engelmann Spruce / Rusty Menziesia / Clasping Twisted-stalk Woodland

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This association has been described primarily as a linear to small-patch type within the northern Rocky Mountains of the U.S. and Canada. It is found from the Clearwater National Forest northward into British Columbia and Alberta, though perhaps it is best developed within northern Idaho and northwestern Montana. Virtually the only site parameter characteristic of all sampled stands is a seasonally high water table, yielding a hygric to subhydric moisture regime. This association is found from mid to upper subalpine sites with elevations ranging from 1340 to 2075 m. It is found on all degrees of slope, in all slope positions and occupying all aspects, with no particular condition considered modal. It should be inferred from the dense canopy of Menziesia ferruginea (accompanied by Rhododendron albiflorum in northern Idaho-eastern Washington) that these sites are also relatively cold. Parent materials run the gamut from sedimentary to intrusive igneous to glacial till and drift with compaction layers. Throughout much of this association''s range a high ash content is present when the type occurs on north- and east-facing slopes. The tree canopy is relatively open, seldom exceeding 60% combined cover and dominated by a combination of Picea engelmannii and Abies lasiocarpa, that can vary between the extremes of nearly monospecific dominance of either species. The undergrowth is dominated by a dense shrub layer, often exceeding 60% canopy cover, in which Menziesia ferruginea is diagnostic and Vaccinium membranaceum approaches 100% constancy. Other shrubs consistently present include Alnus viridis ssp. sinuata, Rubus parviflorus, Ribes lacustre, and Sorbus spp. (predominantly Sorbus sitchensis). The graminoid component is decidedly sparse with only Bromus vulgaris and Luzula glabrata var. hitchcockii being present in more than a third of the sampled stands. Other graminoids present only occasionally, but indicative of a high moisture status, include Cinna latifolia, Calamagrostis canadensis, Vahlodea atropurpurea, and Carex disperma. The forb layer is generally quite species-rich with dominance shifting between any number of species indicative of mesic to relatively wet conditions. The relatively tall forb species considered diagnostic for the type include Athyrium filix-femina (considerably less robust, however, than when occurring in the Thuja plicata alliance), Angelica arguta, Angelica dawsonii, Streptopus amplexifolius, Senecio triangularis, Erigeron peregrinus, Mertensia ciliata, Mertensia paniculata, Aconitum columbianum, Ligusticum canbyi, Veratrum viride, Heracleum maximum, and Trautvetteria caroliniensis; diagnostic forbs of shorter stature include Gymnocarpium dryopteris, Mitella breweri, Mitella pentandra, and Viola glabella.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This association was previously recognized as the Menziesia ferruginea Phase of the Abies lasiocarpa / Streptopus amplexifolius Habitat Type (Cooper et al. 1987). Structurally, compositionally and environmentally this assemblage of stands is sufficiently distinctive to constitute a unique association. It occurs predominantly on upslope positions that are subirrigated and possessing a robust shrub component rather than on subirrigated alluvial bottoms and toeslopes exclusively forb-dominated (or at least lacking the tall-shrub component whose composition is indicative of higher elevation and ostensibly colder sites).

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: The tree canopy is relatively open, seldom exceeding 60% combined cover, and dominated by a combination of Picea engelmannii and Abies lasiocarpa, that can vary between the extremes of nearly monospecific dominance of either species. Other tree species apparently do poorly here as seral associates with only Abies grandis, Pseudotsuga menziesii, Pinus contorta, and Pinus monticola having so much as 10% canopy cover. The undergrowth is dominated by a dense shrub layer, often exceeding 60% canopy cover, in which Menziesia ferruginea is diagnostic and Vaccinium membranaceum approaches 100% constancy. The height of Menziesia exhibits a notable association with regional climates; in northern Idaho where the maritime influence is considerable it grows in excess of 8 feet tall, whereas on the eastern fringes of the northern Rockies (more Continental regime) it does not much exceed 3 feet. Other shrubs consistently present include Alnus viridis ssp. sinuata, Rubus parviflorus, Ribes lacustre, and Sorbus spp. (predominantly Sorbus sitchensis); only infrequently do they constitute as much as 10% cover individually. Rhododendron albiflorum is a common and occasionally high-cover associate in upper elevation stands northward within the association. The graminoid component is decidedly sparse with only Bromus vulgaris and Luzula glabrata var. hitchcockii being present in more than a third of the sampled stands. Other graminoids present only occasionally, but indicative of a high moisture status, include Cinna latifolia, Calamagrostis canadensis, Vahlodea atropurpurea, and Carex disperma. The forb layer is generally quite species-rich with dominance shifting between any number of species indicative of mesic to relatively wet (subhydric) conditions. The relatively tall forb species considered diagnostic for the type include Athyrium filix-femina (considerably less robust, however, than when occurring in the Thuja plicata alliance), Angelica arguta, Angelica dawsonii, Streptopus amplexifolius, Senecio triangularis, Erigeron peregrinus, Mertensia ciliata, Mertensia paniculata, Aconitum columbianum, Ligusticum canbyi, Veratrum viride, Heracleum maximum, and Trautvetteria caroliniensis; diagnostic forbs of shorter stature include Gymnocarpium dryopteris, Mitella breweri, Mitella pentandra, and Viola glabella. Of the foregoing forbs many are regionally restricted. Comprising forbs of high constancy and occasionally dominance are Arnica latifolia (or Arnica cordifolia), Thalictrum occidentale (or Thalictrum fendleri to the south), Galium triflorum, Clintonia uniflora, Tiarella trifoliata, Viola orbiculata, and Xerophyllum tenax (the latter four species found in the association only from central Idaho northward).

Dynamics:  Early-seral conditions within this type have not been documented but, from examination of stand structure of mid-aged stands, it is clear that the seral tree dominants (Abies lasiocarpa and Picea engelmannii) are also the climax dominants. Shade-intolerant (seral) tree species do poorly throughout the sere on these sites, perhaps because the shrub layer responds rapidly following disturbance.

Environmental Description:  This association has been described primarily as a linear to small-patch type within the northern Rocky Mountains of the U.S. and Canada. It is found from the Clearwater National Forest northward into British Columbia and Alberta, though perhaps it is best developed within northern Idaho and northwestern Montana. Virtually the only site parameter characteristic of all sampled stands, and therefore inferred to be the primary ecological driver, is a seasonally high water table, yielding a hygric to subhydric moisture regime. This association is found from mid to upper subalpine sites with elevations ranging from 1340 to 2075 m (4400-6800 feet), though in any Bailey section elevational range is on the order of 475 m (1800 feet) and in a geographic circumscribed area (e.g., Glacier-Waterton International Peace Park) elevation ranges are mostly in the range of 240 to 365 m (800-1200 feet). It is found on all degrees of slope and occupies all aspects, with collecting positions (lower slopes, toeslopes, footslopes, swales) being the norm. It should be inferred from the dense canopy of Menziesia ferruginea (accompanied by Rhododendron albiflorum in northern Idaho-eastern Washington) that these sites are also relatively cold. Effective rooting depths are relatively shallow (average only 36 cm (14 inches) in northern Idaho), which may reflect not only the presence of high water tables and saturated soils but also cold soils. Parent materials run the gamut from sedimentary (both calcareous and noncalcareous) to intrusive igneous (granites of Idaho Batholith) to glacial till and drift with compaction layers. Throughout much of this association''s range, a high ash content is present when the type occurs on north- and east-facing slopes. Adjacent, relatively drier sites usually support ~Abies lasiocarpa - Picea engelmannii / Menziesia ferruginea / Clintonia uniflora Forest (CEGL005893)$$ or ~Abies lasiocarpa - Picea engelmannii / Menziesia ferruginea Forest (CEGL000319)$$.

Geographic Range: This association has been described primarily as a linear or small- to large-patch type within the northern Rocky Mountains of the U.S. and the southwestern corner of the Canadian Rocky Mountains. It is found from the Clearwater National Forest of Idaho northward into Alberta and ostensibly British Columbia, though perhaps it is best developed within northern Idaho and northwestern Montana.

Nations: CA,US

States/Provinces:  AB, ID, MT, WA?




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: G3G4

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Abies lasiocarpa - Picea engelmannii / Menziesia ferruginea / Streptopus amplexifolius Woodland (Hop et al. 2007)
>< Abies lasiocarpa / Rhododendron albiflorum Plant Association (Williams et al. 1995)
> Abies lasiocarpa / Streptopus amplexifolius Habitat Type, Menziesia ferruginea Phase (Hansen et al. 1995)
= Abies lasiocarpa / Streptopus amplexifolius Habitat Type, Menziesia ferruginea Phase (Cooper et al. 1987)
>< Picea engelmannii - Abies lasiocarpa - (Pinus contorta) / Menziesia ferruginea / Arnica cordifolia - Rubus parviflorus Vegetation Type (Achuff et al. 2002)
= Picea engelmannii - Abies lasiocarpa / Menziesia ferruginea - Tiarella unifoliata Habitat Type (Ogilvie 1962)

Concept Author(s): Hop et al. (2007)

Author of Description: S.V. Cooper

Acknowledgements: No Data Available

Version Date: 02-13-04

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