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CEGL007119 Pinus virginiana - Pinus (rigida, echinata) - (Quercus montana) / Vaccinium pallidum Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Virginia Pine - (Pitch Pine, Shortleaf Pine) - (Chestnut Oak) / Blue Ridge Blueberry Forest

Colloquial Name: Appalachian Low-Elevation Mixed Pine / Blue Ridge Blueberry Forest

Hierarchy Level:  Association

Type Concept: This community includes Pinus virginiana-dominated forests of low-elevation ridges and steep upper slopes, occurring primarily in the Appalachian provinces of the eastern United States, from southwestern Virginia, south and west to northern Georgia and northern Alabama. This community occurs on narrow ridges, steep slopes, and other topographic positions with high solar exposure, over shallow, infertile soils. This mainly evergreen forest is often of low stature, with a somewhat open to closed canopy, sparse to very dense shrub cover dominated by ericaceous species, and a sparse herb stratum. Pinus virginiana is the canopy dominant throughout the range of the type. In some parts of the range, other Pinus species may be significant canopy associates, as well as dry-site Quercus species (e.g., Quercus montana, Quercus coccinea). Deciduous species may form a subcanopy or sapling stratum, particularly in areas where fire has been excluded. Common shrub dominants include Vaccinium pallidum, Vaccinium stamineum, Gaylussacia baccata, and Kalmia latifolia. Herbs vary with geography but are typical of infertile, xeric habitats. Some typical herbs in this forest are Baptisia tinctoria, Chimaphila maculata, Dichanthelium commutatum, Epigaea repens, Euphorbia corollata, Galax urceolata, Gaultheria procumbens, Hypoxis hirsuta, Iris verna, Pityopsis graminifolia var. latifolia, Pteridium aquilinum var. latiusculum, and Schizachyrium scoparium.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: In Indiana examples, the substrate is primarily a matrix of acidic siltstone, shale, and sandstone. Rarely are cliffs formed; instead the setting is mostly very steep slopes with high hills and deep ravines. This association also includes vegetation from the transition between the Cumberland Plateau / Southern Ridge and Valley and the Upper East Gulf Coastal Plain in Alabama. Though located in the Coastal Plain, these occurrences are physiographically and floristically similar to this montane association. Related vegetation in Virginia has been treated as several associations within the Central Appalachian Shale Barren and Mountain / Piedmont Acidic Woodlands ecological groups (Fleming and Patterson 2011a).

Early-successional vegetation associated with old fields, old pastures, clearcuts, and burned or eroded areas and dominated by Pinus virginiana is classified as ~Pinus virginiana Ruderal Forest (CEGL002591)$$. Appalachian xeric oak forests with similar floristics, but with a mainly deciduous canopy, are classed in ~Quercus (montana, coccinea) / Kalmia latifolia / (Galax urceolata, Gaultheria procumbens) Forest (CEGL006271)$$. Appalachian shale forests and woodlands with Pinus virginiana occur on steep, shaley slopes and have stunted canopies and sparse herb and shrub strata, characterized by species adapted to shaley substrates. These shale communities are classed in ~Pinus virginiana - Quercus montana Acidic Shale Woodland Alliance (A3312)$$. In 13 plots from the Southern Blue Ridge, species with >54% constancy and at least 10% mean cover are Pinus virginiana, Vaccinium pallidum, Quercus montana, Oxydendrum arboreum, Acer rubrum, Quercus coccinea, Nyssa sylvatica, Pinus rigida, Gaylussacia baccata, and Gaylussacia ursina.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: This community is a needle-leaved evergreen forest with a usually somewhat open (occasionally closed) canopy. The canopy is typically short (<20 m) with tree height and canopy cover decreasing with increasing severity of the microsite. A deciduous subcanopy may be present. The shrub layers can be sparse but are more often dense to very dense and are composed of tall and short shrubs, predominantly ericaceous species. Herb cover is sparse, and leaf litter often dominates the ground layer. Pinus virginiana is the canopy dominant throughout the range of the type. In the Southern Appalachians and Southern Ridge and Valley it may occur with mixes of Pinus rigida, Pinus echinata, or Pinus strobus. Within its range, Pinus pungens may be present as a very minor component. Regeneration of Pinus virginiana is concentrated along cliff edges and tends to drop off inward from the edge. Small stems of Quercus montana (= Quercus prinus), Quercus coccinea, Acer rubrum, Nyssa sylvatica, and Oxydendrum arboreum are common in the subcanopy and sapling strata and may occur in the canopy as well. In the Southern Blue Ridge/Piedmont and Southern Blue Ridge/Ridge and Valley transition regions, Quercus marilandica, Quercus falcata, and Quercus stellata can be deciduous components. Common shrub dominants include Vaccinium pallidum, Vaccinium stamineum, Gaylussacia baccata, and Kalmia latifolia. Other typical shrubs can include Gaylussacia ursina, Kalmia latifolia, Sassafras albidum, and Vaccinium hirsutum (southwestern North Carolina and southeastern Tennessee only). Smilax glauca and Smilax rotundifolia can be common vines. In the sparse herb layer, characteristic species from the Southern Blue Ridge and Southern Ridge and Valley include Baptisia tinctoria, Chimaphila maculata, Dichanthelium commutatum, Danthonia spicata, Epigaea repens, Euphorbia corollata, Galax urceolata, Gaultheria procumbens, Hypoxis hirsuta, Iris verna, Pityopsis graminifolia var. latifolia, Pteridium aquilinum var. latiusculum, and Schizachyrium scoparium. Typical herbs from examples in the western portion of the range (Interior Low Plateau) include Antennaria plantaginifolia, Antennaria solitaria, Carex albicans var. albicans (= Carex artitecta), Danthonia spicata, Dichanthelium dichotomum, Lespedeza violacea (= Lespedeza intermedia), Hieracium gronovii, Hieracium venosum, Krigia biflora, Solidago erecta, and Tephrosia virginiana (M. Homoya pers. comm. 1999). In some of these examples Opuntia humifusa, Calamagrostis porteri ssp. insperata, and Solidago squarrosa may occur locally. Nonvascular plants have not been documented rangewide; West Virginia examples have identified lichen species including Cladonia caroliniana, Cladonia squamosa, Cladonia rangiferina, Cladonia pyxidata, Umbilicaria mammulata, Lasallia papulosa, Lasallia pensylvanica, Xanthoparmelia plittii, Xanthoparmelia conspersa, and Flavoparmelia caperata, and moss species including Leucobryum glaucum, Leucobryum albidum, Polytrichum juniperinum, Dicranum scoparium, Dicranum spurium, Dicranum condensatum, Hypnum imponens, and Thuidium delicatulum. In Indiana''s Knobstone Escarpment, this community provides habitat for that state''s only endemic vascular plant, Penstemon deamii.

Dynamics:  This xeric, evergreen forest community will be maintained on sites where local soil conditions and topographic extremes function to retard hardwood invasion. Although Pinus virginiana can become established by seed following fire, the trees are thin-barked and are easily killed by fire. Repeated burning is likely to select for Pinus rigida at the expense of Pinus virginiana. Infestations of southern pine beetle (Dendroctonus frontalis) can cause mortality of canopy trees. Examples affected by southern pine beetle in the Great Smoky Mountains can have up to 80-90% standing dead pine. This forest is also susceptible to damage by fire and has been found to be particularly sensitive to ozone, which causes foliar damage of Pinus virginiana (Burns and Honkala 1990a).

This community may occur as early- to late-successional stands. On sites where local soil conditions or topographic extremes retard succession and maintain pine dominance indefinitely, this community is considered a topo-edaphic climax. Other occurrences may be successional to forests with mixed deciduous-evergreen canopies dominated by mixtures of Pinus (pungens, rigida, virginiana, echinata) and Quercus (coccinea, montana) species.

Throughout most of its range, this community occurs as linear features along ridgetops and may be adjacent to or grade into xeric forests dominated by Quercus coccinea or Quercus montana or more mesic forests dominated by Quercus alba, Quercus rubra, Quercus velutina, Carya glabra, and Carya alba. In the Interior Low Plateau, individual stands can be small in size, occurring in a matrix of Quercus montana or Quercus montana - Quercus alba forest (e.g., ~Quercus montana / Smilax spp. Forest (CEGL005022)$$ or ~Quercus montana - Quercus (alba, coccinea) / Viburnum acerifolium - (Kalmia latifolia) Forest (CEGL005023)$$), but in more edaphically extreme circumstances.

Environmental Description:  Stands of this forest occur on narrow ridges and knobs, steep upper slopes, bluff and cliff tops, and other exposed sites throughout the range of the type. The community is found primarily on south-, southeast- or southwest-facing aspects on excessively drained, shallow soils. In the Blue Ridge Escarpment region, the western margin of the Blue Ridge, and west into the Ridge and Valley and Cumberland Mountains, this xeric forest occurs on convex slopes and ridges below 850 m (2800 feet) elevation, over soils classed as Inceptisols, typically Lithic Dystrochrepts originating from sandstone, shale and other noncalcareous parent material. Occurrences in rugged parts of the western Piedmont are also likely. Its environmental situation in the western Alleghenies is not known. In the Interior Low Plateau of Kentucky, Tennessee, and Indiana, this association occurs in edaphically extreme situations, including bluff tops and narrow ridges in thin soils weathered from relatively acidic caprocks with southern and western aspects, as well as other similar slopes, over cherty limestone, siltstones, sandstones, and shales. In particular, in the Knobstone Escarpment Subsection (a few Indiana counties just north of Louisville, Kentucky), it occurs in gladelike situations on steep slopes with thin soils. Along the edges of cliff tops, there is usually a narrow zone of exposed bedrock pavement and patches of very shallow soil, but soils become progressively deeper back from the cliff edge. At least in West Virginia, portions of the stands along the cliff edge are likely to be edaphic climax communities, but farther back from the edge, they are likely to be successional following fire.

Geographic Range: This community occurs primarily in the Appalachian region of the United States, ranging from southwestern Virginia, south and west through the Ridge and Valley, Blue Ridge, and Cumberland Plateau to northern Georgia and Alabama, extending westward to scattered areas in the Interior Low Plateau and eastward into the upper Piedmont. It is recorded from the states of Georgia, North Carolina, South Carolina, Tennessee, Kentucky, Pennsylvania, Indiana, Ohio, Maryland, and West Virginia.

Nations: US

States/Provinces:  AL, GA, IN, KY, NC, OH, SC, TN, WV




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: G3

Greasons: No Data Available


Concept Lineage: merged. CEGL006224 and CEGL007082 merged with CEGL006061 which was subsequently merged into CEGL007119. CEGL007538 merged with CEGL005040 (which was subsequently merged into CEGL007119).

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Pinus virginiana - (Quercus spp.) / Nyssa sylvatica / Gaultheria procumbens forest (Vanderhorst 2002b)
= Pinus virginiana - Pinus (rigida, echinata) - (Quercus prinus) / Vaccinium pallidum Forest (Faber-Langendoen 2001)
? Pinus virginiana - Quercus prinus - Nyssa sylvatica Forest (Walton et al. 1997)
< IA7c. Xeric Virginia Pine Ridge Forest (Allard 1990)
< Oligotrophic Forest (Rawinski 1992)
< Virginia Pine - Oak: 78 (Eyre 1980)
< Virginia Pine Type (Schmalzer and DeSelm 1982)
< Virginia Pine: 79 (Eyre 1980)
? Virginia pine forest (CAP pers. comm. 1998)

Concept Author(s): K.D. Patterson

Author of Description: K.D. Patterson, R. White and S.C. Gawler

Acknowledgements: No Data Available

Version Date: 04-01-10

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