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CEGL006226 Tsuga canadensis - Betula alleghaniensis / Ilex verticillata / Sphagnum spp. Swamp Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Eastern Hemlock - Yellow Birch / Common Winterberry / Peatmoss species Swamp Forest

Colloquial Name: Hemlock - Hardwood Swamp Forest

Hierarchy Level:  Association

Type Concept: These hemlock-hardwood swamps are common throughout most of New England, New York and Pennsylvania, mostly in glaciated areas south of spruce-fir forest regions. Most are mixed-canopy wetland forests, but in some may be strongly coniferous. They occur in poorly drained basins over bedrock or compacted till. "Pocket swamps," small isolated basins in upland forests, are one setting, and these swamps also occur adjacent to streams and lakes in larger basins. The acidic, organic soils remain saturated for most or all of the growing season; they may partially dry out, particularly in smaller basins. Canopy closure is nearly complete, and shrubs are sparse. The herbaceous layer may be well-developed, with ferns especially characteristic. Bryophyte cover varies, but is usually extensive. Tsuga canadensis is either canopy dominant or is mixed with other trees including Acer rubrum, Betula alleghaniensis, Pinus strobus, Nyssa sylvatica, and Fraxinus nigra. Scattered shrubs include Vaccinium corymbosum, Ilex verticillata, Lyonia ligustrina, Ilex mucronata, and Viburnum nudum var. cassinoides. Osmunda cinnamomea, Osmunda regalis, and Onoclea sensibilis are prominent ferns; associated herbs include Carex folliculata, Carex trisperma, Carex disperma, Maianthemum canadense, Coptis trifolia, Dryopteris cristata, Rubus pubescens, and Mitchella repens. Bryophytes include Sphagnum girgensohnii, Sphagnum palustre, and other Sphagnum species, as well as Pleurozium schreberi and Bazzania trilobata. These forests lack the species of more southerly affinity, such as Rhododendron maximum or Liriodendron tulipifera, that characterize hemlock swamps to the south, which are covered by ~Tsuga canadensis / Rhododendron maximum / Sphagnum spp. Swamp Forest (CEGL006279)$$. Mixed expression of these swamps may be similar in canopy composition to ~Betula alleghaniensis - Acer rubrum - (Tsuga canadensis, Abies balsamea) / Osmunda cinnamomea Swamp Forest (CEGL006380)$$; those are mineral-soil wetlands in seepage-influenced areas, typically at wetland-upland ecotones, and usually occur on slopes rather than in basins. ~Acer rubrum - Nyssa sylvatica - Betula alleghaniensis / Sphagnum spp. Swamp Forest (CEGL006014)$$ is ecologically similar but is more strongly deciduous, with limited hemlock.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: No Data Available

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: Canopy closure is nearly complete, and shrubs are sparse. The herbaceous layer may be well-developed, with ferns especially characteristic. Bryophyte cover varies, but is usually extensive. Tsuga canadensis is either canopy dominant or is mixed with other trees including Acer rubrum, Betula alleghaniensis, Pinus strobus, Nyssa sylvatica, and Fraxinus nigra. Scattered shrubs include Vaccinium corymbosum, Ilex verticillata, Lyonia ligustrina, Ilex mucronata (= Nemopanthus mucronatus), and Viburnum nudum var. cassinoides. Osmunda cinnamomea, Osmunda regalis, and Onoclea sensibilis are prominent ferns; associated herbs include Carex folliculata, Carex trisperma, Carex disperma, Maianthemum canadense, Coptis trifolia, Dryopteris cristata, Rubus pubescens, and Mitchella repens. Bryophytes include Sphagnum girgensohnii, Sphagnum palustre, and other Sphagnum species, as well as Pleurozium schreberi and Bazzania trilobata. These forests lack the species of more southerly affinity, such as Rhododendron maximum or Liriodendron tulipifera, that characterize hemlock swamps to the south.

Dynamics:  No Data Available

Environmental Description:  These hemlock-hardwood swamps are common throughout most of New England, New York and Pennsylvania, mostly in glaciated areas south of spruce-fir forest regions. Most are mixed-canopy wetland forests, but in some may be strongly coniferous. They occur in poorly drained basins over bedrock or compacted till. "Pocket swamps," small isolated basins in upland forests, are one setting, and these swamps also occur adjacent to streams and lakes in larger basins. The acidic, organic soils remain saturated for most or all of the growing season; they may partially dry out, particularly in smaller basins.

Geographic Range: This community occurs in glaciated areas ranging from New Jersey and Pennsylvania north through New York and New England to New Brunswick and possibly Nova Scotia, Canada.

Nations: CA,US

States/Provinces:  CT, MA, ME, NB, NH, NJ, NS?, NY, PA, RI, VT




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: G5

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: < Hardwood-Conifer Swamp (Breden 1989)
< Hemlock - Yellow Birch: 24 (Eyre 1980)
< Inland Atlantic white cedar swamp (Reschke 1990)
< Northern New England basin swamp (Rawinski 1984a)

Concept Author(s): J. Lundgren

Author of Description: J. Lundgren and E. Largay

Acknowledgements: No Data Available

Version Date: 08-23-06

  • Breden, T. F. 1989. A preliminary natural community classification for New Jersey. Pages 157-191 in: E. F. Karlin, editor. New Jersey''s rare and endangered plants and animals. Institute for Environmental Studies, Ramapo College, Mahwah, NJ. 280 pp.
  • Breden, T. F., Y. R. Alger, K. S. Walz, and A. G. Windisch. 2001. Classification of vegetation communities of New Jersey: Second iteration. Association for Biodiversity Information and New Jersey Natural Heritage Program, Office of Natural Lands Management, Division of Parks and Forestry, New Jersey Department of Environmental Protection, Trenton.
  • Davis, T. 2011b. Pennsylvania Natural Heritage Program. Hemlock - Mixed Hardwood Palustrine Forest Factsheet. [http://www.naturalheritage.state.pa.us/Community.aspx?=16029] (accessed January 31, 2012)
  • Eastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boston, MA.
  • Edinger, G. J., D. J. Evans, S. Gebauer, T. G. Howard, D. M. Hunt, and A. M. Olivero, editors. 2014a. Ecological communities of New York state. Second edition. A revised and expanded edition of Carol Reschke''s ecological communities of New York state. New York Natural Heritage Program, New York State Department of Environmental Conservation, Albany, NY.
  • Enser, R. W., and J. A. Lundgren. 2006. Natural communities of Rhode Island. A joint project of the Rhode Island Department of Environmental Management Natural Heritage Program and The Nature Conservancy of Rhode Island. Rhode Island Natural History Survey, Kingston. 40 pp. [www.rinhs.org]
  • Eyre, F. H., editor. 1980. Forest cover types of the United States and Canada. Society of American Foresters, Washington, DC. 148 pp.
  • Fike, J. 1999. Terrestrial and palustrine plant communities of Pennsylvania. Pennsylvania Natural Diversity Inventory. Pennsylvania Department of Conservation and Recreation, Bureau of Forestry, Harrisburg, PA. 86 pp.
  • Gawler, S. C. 2002. Natural landscapes of Maine: A guide to vegetated natural communities and ecosystems. Maine Natural Areas Program, Department of Conservation, Augusta, ME.
  • Gawler, S. C., and A. Cutko. 2010. Natural landscapes of Maine: A classification of vegetated natural communities and ecosystems. Maine Natural Areas Program, Department of Conservation, Augusta.
  • Lundgren, J. 2000. Lower New England - Northern Piedmont Ecoregion Forest Classification. The Nature Conservancy, Conservation Science, Boston, MA. 72 pp.
  • Messier, S. N. 1980. The plant communities of the acid wetlands of northwestern Connecticut. M.S. thesis, University of Connecticut, Storrs. 96 pp.
  • Metzler, K., and J. Barrett. 2006. The vegetation of Connecticut: A preliminary classification. State Geological and Natural History Survey, Report of Investigations No. 12. Connecticut Natural Diversity Database, Hartford, CT.
  • Perles, S. J., G. S. Podniesinski, M. Furedi, B. A. Eichelberger, A. Feldmann, G. Edinger, E. Eastman, and L. A. Sneddon. 2008. Vegetation classification and mapping at Upper Delaware Scenic and Recreational River. Technical Report NPS/NER/NRTR--2008/133. National Park Service, Philadelphia, PA. 370 pp.
  • Rawinski, T. 1984a. Natural community description abstract - southern New England calcareous seepage swamp. Unpublished report. The Nature Conservancy, Boston, MA. 6 pp.
  • Reschke, C. 1990. Ecological communities of New York State. New York Natural Heritage Program, New York State Department of Environmental Conservation, Latham, NY. 96 pp.
  • Sechler, F. C., G. J. Edinger, T. G. Howard, J. J. Schmid, E. Eastman, E. Largay, L. A. Sneddon, C. Lea, and J. Von Loh. 2014. Vegetation classification and mapping at Roosevelt-Vanderbilt National Historic Sites, New York. Natural Resource Technical Report NPS/NETN/NRTR--2014/873, National Park Service, Fort Collins, CO. 392 pp.
  • Sperduto, D. D., and W. F. Nichols. 2004. Natural communities of New Hampshire: A guide and classification. New Hampshire Natural Heritage Bureau, DRED Division of Forests and Lands, Concord. 242 pp.
  • Swain, P. C., and J. B. Kearsley. 2014. Classification of the natural communities of Massachusetts. Version 2.0. Natural Heritage & Endangered Species Program, Massachusetts Division of Fisheries and Wildlife. Westborough, MA. [http://www.mass.gov/nhesp/http://www.mass.gov/eea/agencies/dfg/dfw/natural-heritage/natural-communities/classification-of-natural-communities.html]
  • Thompson, E. H., and E. R. Sorenson. 2005. Wetland, woodland, wildland: A guide to the natural communities of Vermont. The Nature Conservancy and the Vermont Department of Fish and Wildlife. University Press of New England, Hanover, NH. 456 pp.
  • Zimmerman, E. A., T. Davis, M. A. Furedi, B. Eichelberger, J. McPherson, S. Seymour, G. Podniesinski, N. Dewar, and J. Wagner, editors. 2012. Terrestrial and palustrine plant communities of Pennsylvania. Pennsylvania Natural Heritage Program, Pennsylvania Department of Conservation and Natural Resources, Harrisburg. [http://www.naturalheritage.state.pa.us/Communities.aspx]