Print Report

CEGL006376 Ilex opaca / Morella pensylvanica Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: American Holly / Northern Bayberry Forest

Colloquial Name: Maritime Holly Forest

Hierarchy Level:  Association

Type Concept: This type represents maritime holly forest of the lee sides of backdunes in New Jersey and Long Island, New York. It occurs in low areas behind backdunes where it is protected from overwash and salt spray. Substrate is sandy loam over coarse sand. The dominant tree is Ilex opaca. Other canopy associates can include Amelanchier canadensis, Amelanchier laevis, Acer rubrum, Celtis occidentalis, Sassafras albidum, Quercus stellata, Quercus velutina, and Prunus serotina. Characteristic shrubs include Morella pensylvanica, Gaylussacia baccata, Rhus copallinum, Vaccinium corymbosum. Sambucus canadensis, Viburnum dentatum, and Toxicodendron vernix can occur infrequently. Vines are particularly abundant, and include Toxicodendron radicans, Smilax rotundifolia, Parthenocissus quinquefolia, and Vitis spp. The herbaceous layer is sparsely to moderately developed and includes Aralia nudicaulis, Maianthemum stellatum, Maianthemum canadense, Maianthemum racemosum, Carex pensylvanica, and Cypripedium acaule. In locally wetter, boggy areas within the forest Nyssa sylvatica, Rhododendron viscosum, Vaccinium corymbosum, Photinia melanocarpa, Osmunda cinnamomea, Osmunda regalis, and Thelypteris palustris occur abundantly.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Occurrences in New York and New Jersey are on the order of 50+ acres in size.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: The dominant tree is Ilex opaca; stems can be 300 years old. Other canopy associates can include Amelanchier canadensis, Amelanchier laevis, Acer rubrum, Celtis occidentalis, Sassafras albidum, Quercus stellata, Quercus velutina and Prunus serotina. Characteristic shrubs include Morella pensylvanica (= Myrica pensylvanica), Gaylussacia baccata, Rhus copallinum, and Vaccinium corymbosum. Sambucus canadensis, Viburnum dentatum, and Toxicodendron vernix can occur infrequently. Vines are particularly abundant and include Toxicodendron radicans, Smilax rotundifolia, Parthenocissus quinquefolia, and Vitis spp. The herbaceous layer is sparsely to moderately developed and includes Aralia nudicaulis, Maianthemum stellatum (= Smilacina stellata), Maianthemum racemosum (= Smilacina racemosa), and Maianthemum canadense. In locally wetter, boggy areas within the forest Nyssa sylvatica, Rhododendron viscosum, Vaccinium corymbosum, Photinia melanocarpa, Osmunda cinnamomea, Osmunda regalis, and Thelypteris palustris occur abundantly.

Dynamics:  This is a late-successional expression of wet-mesic forest that develops behind protected maritime dunes. Reproduction is vegetative, not via seeds (Art 1976). Deer browsing affects the shrub and herb layers of sunken forests more than canopy composition decreasing overall species diversity (Art 1992). On Sandy Hook, in New Jersey, the holly forest occurs just inland from a salt marsh.

Environmental Description:  This sunken forest occurs in hollows leeward of maritime backdunes where they are protected from tidal overwash and salt spray except during severe storms and hurricanes. Substrate is generally sandy loam over coarse sand with local pockets of peat.

Geographic Range: Currently only known from New Jersey and Long Island, New York.

Nations: US

States/Provinces:  NJ, NY




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: G1

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Maple-holly forest (Chrysler 1930)
= Sunken Forest (Greller 1977)
= Sunken Forest (Art 1976)
? Sunken Forest (Stalter 1979)

Concept Author(s): D. Hunt (NYNHP)

Author of Description: S.L. Neid and E. Largay

Acknowledgements: No Data Available

Version Date: 03-26-07

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  • Art, H. W. 1987. Patterns of community dynamics in the Sunken Forest: 1967 to 1985 and 1985 to 1986. National Park Service, North Atlantic Regional Office. 66 pp.
  • Art, H. W. 1992. The impacts of Hurricane Gloria on deer and trails, the Sunken Forest, Fire Island National Seashore, Fire Island, NY. National Park Service, North Atlantic region, Boston, MA 168 pp.
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