Print Report

CEGL000476 Thuja plicata / Gymnocarpium dryopteris Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Western Red-cedar / Western Oakfern Forest

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This Thuja plicata forest association is known from the northern Rocky Mountains of British Columbia, northern Idaho and northwestern Montana. It represents one of the driest Thuja plicata riparian communities, and the driest Thuja type to have a fern layer. It occurs at elevations ranging from 975 to 1370 m (3200-4500 feet). Typical locations include slopes or benches along major mountain streams. Slopes are moderate to steep, and stands are generally located on mid to lower slopes. Parent materials are usually quartzite, sandstone, or schist, often mixed or overlain with volcanic ash. Soils are gravelly loams to silty clay loam to silt. Stands are subirrigated yet well-drained. Duff layers average 8 cm in depth. Late-seral stands have closed canopies and are dominated by Thuja plicata. Stands not in a late-seral stage are more heterogeneous, with a mix of conifers in addition to Thuja. Seral species persisting in these stands include Abies grandis as the most abundant and common. Others include Tsuga heterophylla, Picea engelmannii (or Picea x albertiana), Pinus monticola, Larix occidentalis, and Pseudotsuga menziesii. The shrub layer is typically limited to 10-20% total cover. Highly constant species include Acer glabrum, Lonicera utahensis, Linnaea borealis, Rubus parviflorus, and Taxus brevifolia. The herbaceous layer has mostly ferns and forb species with few to no grasses. Gymnocarpium dryopteris is diagnostic for this type, with at least 1% cover, but is often much more abundant. Athyrium filix-femina or Adiantum pedatum are often present, but with low cover or only on moist microsites. Other highly constant mesic forbs include Clintonia uniflora, Tiarella trifoliata, Coptis occidentalis, Osmorhiza berteroi, Prosartes hookeri, Aralia nudicaulis, and Maianthemum stellatum.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This community, in regard to subcanopy and undergrowth composition, defining abiotic parameters and geographic distribution is virtually indistinguishable from ~Tsuga heterophylla / Gymnocarpium dryopteris Forest (CEGL000494)$$. The difference between Tsuga and Thuja dominance of the upper canopy is quite probably related to historical accident, past disturbance events and subsequent successional patterns. Despite the very long fire-return intervals (200-500 years for stand-replacing fire), the longevity of both species, especially Thuja, argues for considering the dual designation (Thuja plicata - Tsuga heterophylla) as appropriate (i.e., combining these two associations into one); if one or the other species responds to disturbance by attaining canopy dominance, it is unlikely the non-dominant species will become dominant in the average fire-free interval. Others have recognized that quite probably a dominance continuum exists between these two species by naming at least 8 plant associations with the dual designation (Tsuga heterophylla - Thuja plicata /_).

Arguing against this approach is the approach being taken by the USNVC placing emphasis on existing vegetation composition, and it is indisputable that either climax tree species can be strongly dominant with the other species only occurring in the reproductive layers, if at all. In northern Idaho, where these species are sympatric over an extensive range, there are drainages where one or the other species is present and its complement is not (Daubenmire and Daubenmire 1968); this phenomenon has never been satisfactorily explained and again argues for recognizing separate Thuja plicata and Tsuga heterophylla types.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: Late-seral stands are dominated by Thuja plicata (3-90%). Stands are often not in a late-seral stage and can have several other conifer species. Seral species persisting in these stands include Abies grandis as the most abundant and common with 1-60% cover. Others include Tsuga heterophylla, Picea engelmannii (or Picea x albertiana (= Picea engelmannii x glauca)), Pinus monticola, Larix occidentalis, and Pseudotsuga menziesii, usually present with 1-10% cover. This is the wettest of the closed-canopy Thuja plicata forest types to have an appreciable shrub component (Cooper et al. 1987), with the exception of mature stands of ~Thuja plicata - Tsuga heterophylla / Oplopanax horridus Rocky Mountain Swamp Forest (CEGL000479)$$. The shrub layer is typically limited to 10-20% total cover. Highly constant species include Acer glabrum, Lonicera utahensis, Linnaea borealis, Rubus parviflorus, and Taxus brevifolia. Others occasionally present include Menziesia ferruginea, Vaccinium membranaceum, Cornus sericea, and Oplopanax horridus (but with less than 5% cover). The herbaceous layer has mostly ferns and forb species with few to no grasses. Gymnocarpium dryopteris is diagnostic for this type, with at least 1% cover, but is often much more abundant. Athyrium filix-femina or Adiantum pedatum are often present, but with low cover or only on moist microsites. Other highly constant mesic forbs include Clintonia uniflora, Tiarella trifoliata, Coptis occidentalis, Osmorhiza berteroi, Prosartes hookeri (= Disporum hookeri), Aralia nudicaulis, and Maianthemum stellatum.

Dynamics:  Fire-return intervals for this and related mesic to hygric associations are estimated to be on the order of 200 to 400 years. The moisture regime of these sites is such that the climax species (Thuja plicata and Tsuga heterophylla) are able to rapidly recolonize disturbed sites, even those experiencing stand-replacing fire. Thus these sites may be occupied by early-seral forest as well as late-successional and climax stages. Though highly reduced in abundance and occurring only in microsite positions, mesophytic forbs consistently survive fire (stand-replacing and surface fires) and logging; once a shrub/tree canopy is re-established they flourish to characterize the undergrowth with their lushness and diversity. At the canopy closure stage of succession the undergrowth may be reduced to scattered forbs; often Gymnocarpium dryopteris is the most conspicuous, though its stature may be reduced in microsite positions.

Environmental Description:  This association is one of the driest Thuja plicata riparian communities, and the driest Thuja type to have a fern layer. It occurs at elevations ranging from 975 to 1370 m (3200-4500 feet). Typical locations include slopes or benches along major mountain streams. Slopes are moderate to steep, and stands are generally located on mid to lower slopes. Parent materials are usually quartzite, sandstone, or schist, often mixed or overlain with volcanic ash (Hansen et al. 1995). Soils are gravelly loams to silty clay loam to silt. Stands are subirrigated yet well-drained. Duff layers average 8 cm in depth.

Geographic Range: This association is known from areas of inland maritime climate of the northern Rocky Mountains, in British Columbia, northern Idaho and northwestern Montana.

Nations: CA?,US

States/Provinces:  BC?, ID, MT, WA




Confidence Level: Low - Poorly Documented

Confidence Level Comments: No Data Available

Grank: G3

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: >< Thuja plicata / Asarum caudatum Habitat Type, Gymnocarpium dryopteris Phase (Steele et al. 1976)
>< Thuja plicata / Clintonia uniflora Habitat Type, Aralia nudicaulis Phase (Pfister et al. 1977) [wetter stands of this habitat type.]
>< Thuja plicata / Clintonia uniflora Plant Association (Williams et al. 1995) [Small portion of this type where Thuja is dominant, and Gymnocarpium is present (at least 5% cover) is included here.]
< Thuja plicata / Gymnocarpium dryopteris Habitat Type (Cooper et al. 1987)
< Tsuga heterophylla / Gymnocarpium dryopteris Habitat Type (Cooper et al. 1987) [includes stands where Thuja has >25% relative cover in the canopy, and Tsuga has less are included here.]
>< Tsuga heterophylla / Gymnocarpium dryopteris Plant Association (Williams et al. 1995) [Small portion of this type where Thuja is dominant, and Gymnocarpium is present (at least 5% cover) is included here.]
< Tsuga heterophylla / Pachistima myrsinites Habitat Type (Daubenmire and Daubenmire 1968) [includes stands where Thuja is predominant, and Gymnocarpium is present (at least 1% cover).]

Concept Author(s): Western Ecology Group

Author of Description: G. Kittel

Acknowledgements: No Data Available

Version Date: 04-15-04

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