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CEGL002378 Pinus strobus - (Pinus resinosa) Driftless Bluff Forest
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: Eastern White Pine - (Red Pine) Driftless Bluff Forest
Colloquial Name: White Pine - Red Pine Driftless Bluff Forest
Hierarchy Level: Association
Type Concept: This pine bluff forest occurs on steep sandstone or, rarely, limestone bluffs in the Driftless Area of the upper midwestern United States. Stands occur in small patches 0.1-2 ha in size. The soils are shallow, poorly developed, and often acidic. Runoff and leaching are severe due to the slope and the porous nature of the bedrock. Pinus strobus typically dominates the overstory, with Pinus resinosa and Tsuga canadensis either present or locally dominant. Other hardwood associates include Betula papyrifera, Populus tremuloides, Quercus alba, Quercus velutina, Ostrya virginiana. The composition of the understory is variable but typically contains herbs and shrubs with more northerly distributions, such as Diervilla lonicera, Vaccinium myrtilloides, Eurybia macrophylla, Chimaphila umbellata, Gaultheria procumbens, Lycopodium obscurum, Mitchella repens, Pyrola americana, and Trientalis borealis, as well as some prairie species, including Amorpha canescens, Coreopsis palmata, and Dodecatheon meadia. This community is a relict, thought to have existed in the Driftless Area since at least the Wisconsin glaciation.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: This type should be compared to the more moist ~Pinus strobus - Abies balsamea - Betula alleghaniensis Driftless Forest (CEGL002111)$$.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: The vegetation is a closed to open canopy of conifer or mixed conifer-hardwood trees. The canopy is typically dominated by Pinus strobus. Occasionally Tsuga canadensis is mixed with the pine, or may dominate in certain areas, typically the most steep areas. Some of the stands examined by McIntosh (1950) had pine seedlings and saplings in the understory, while others were dominated by young hardwoods, such as Betula papyrifera, Populus tremuloides, Quercus alba, Quercus velutina, Ostrya virginiana, or Tilia americana. Less common conifer associates include Juniperus virginiana and Pinus banksiana. Many of the herbs and shrubs of the northern pine forests are present in these forests. Typical shrub associates include Diervilla lonicera and Vaccinium myrtilloides. Less common shrubs include Acer spicatum, Cornus canadensis, Corylus cornuta, Prunus pensylvanica, and Prunus virginiana. Typical herbs include Maianthemum canadense. Less common herb and dwarf-shrub associates may include Eurybia macrophylla (= Aster macrophyllus), Chimaphila umbellata, Gaultheria procumbens, Lycopodium obscurum, Mitchella repens, Pyrola americana (= Pyrola rotundifolia), and Trientalis borealis. Other members of the understory are found more commonly on prairies, including Amorpha canescens, Coreopsis palmata, and Dodecatheon meadia. Species typical of more southern hardwood stands include Arisaema triphyllum and Sanguinaria canadensis.
Variability in species composition in the shrub and herbaceous layers is high. This is due to the small, isolated nature of stands. No quantitative data were found on species variability, but both McIntosh (1950) and Curtis (1959) stated that it was high. This variability is due to at least two factors, both of which have to do with the small and isolated nature of examples of this community. One reason is that if a population of a northern species is extirpated from one site, it is unlikely to recolonize due to the distance to another seed source (another stand of this community) (McIntosh 1950). The second reason is that the makeup of the understory is also dependent on that of the surrounding vegetation. Because this community rarely exceeds 2 ha, some species from adjacent communities can easily invade. The identity of these species varies depending on the composition of the nearby communities.
Variability in species composition in the shrub and herbaceous layers is high. This is due to the small, isolated nature of stands. No quantitative data were found on species variability, but both McIntosh (1950) and Curtis (1959) stated that it was high. This variability is due to at least two factors, both of which have to do with the small and isolated nature of examples of this community. One reason is that if a population of a northern species is extirpated from one site, it is unlikely to recolonize due to the distance to another seed source (another stand of this community) (McIntosh 1950). The second reason is that the makeup of the understory is also dependent on that of the surrounding vegetation. Because this community rarely exceeds 2 ha, some species from adjacent communities can easily invade. The identity of these species varies depending on the composition of the nearby communities.
Dynamics: This community is apparently stable in most of its current sites. However, treefall is relatively common in this community due to the steep slopes and shallow soils. In some areas hardwoods more typical of the surrounding forest dominate the understory and may eventually replace the pines in the overstory. On most sites the conditions favor the maintenance of the community. Given the right conditions, lack of fire and no grazing, the pines have even invaded adjacent hardwood forest in a few instances.
Environmental Description: This community occurs on sloping or level sandstone or, rarely, on quartzite bluffs. McIntosh (1950) studied 22 stands of this type and found them on all aspects except southeast and all were on St. Peter or Franconia sandstone of Cambrian or Ordovician age. In Minnesota, this community has been reported on limestone (H. Dunevitz pers. comm.). St. Peter sandstone is very friable and porous, allowing leaching and rapid runoff. It is composed of silica with small amounts of iron and clay. Franconia sandstone is coarse-grained, loosely cemented together, and usually shaley (McIntosh 1950). Soils are sandy, shallow to very shallow, usually acidic, and excessively drained. The most common site is a waterworn cliff with the undercutting stream still present at the cliff base. Considerable light penetrates to the forest floor even though the trees are close together.
The steep slopes combined with the shallow soils prevent other trees from becoming prominent members of the community. The dry, cooler microclimate afforded these pine relicts by the slope and aspect on which they occur also favors their maintenance. McIntosh (1950) states that northerly aspects are cooler and westerly aspects are drier in his study area. Both of these conditions favor conifers over broadleaf trees. A site that faced the warmer, wetter direction (southeast) would be less likely to support the vegetation of this type.
The steep slopes combined with the shallow soils prevent other trees from becoming prominent members of the community. The dry, cooler microclimate afforded these pine relicts by the slope and aspect on which they occur also favors their maintenance. McIntosh (1950) states that northerly aspects are cooler and westerly aspects are drier in his study area. Both of these conditions favor conifers over broadleaf trees. A site that faced the warmer, wetter direction (southeast) would be less likely to support the vegetation of this type.
Geographic Range: This community occurs in many places in the Driftless Area of southwestern Wisconsin, and also in a few places in the area of Farmdale glaciation. It is also found in southeastern Minnesota and in very small occurrences in northeastern Iowa.
Nations: US
States/Provinces: IA, MN?, WI
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.686787
Confidence Level: Moderate
Confidence Level Comments: No Data Available
Grank: G2G3
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.2 Cool Temperate Forest & Woodland Formation | F008 | 1.B.2 |
Division | 1.B.2.Na Eastern North American Forest & Woodland Division | D008 | 1.B.2.Na |
Macrogroup | 1.B.2.Na.4 White Oak - Bur Oak - Shagbark Hickory Forest, Woodland & Savanna Macrogroup | M012 | 1.B.2.Na.4 |
Group | 1.B.2.Na.4.b North-Central Oak - Hickory Forest & Woodland Group | G649 | 1.B.2.Na.4.b |
Alliance | A3237 Eastern White Pine - Oak species Driftless Forest & Woodland Alliance | A3237 | 1.B.2.Na.4.b |
Association | CEGL002378 Eastern White Pine - (Red Pine) Driftless Bluff Forest | CEGL002378 | 1.B.2.Na.4.b |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: = Pinus strobus - (Pinus resinosa) Driftless Bluff Forest (Faber-Langendoen 2001) [Northern Dry-Mesic Forest White Pine-Red Pine Relict]
= White Pine Forest (Southeast Section) (MNNHP 1993)
= White Pine Forest (Southeast Section) (MNNHP 1993)
- Cahayla-Wynne, R., and D. C. Glenn-Lewin. 1978. The forest vegetation of the Driftless Area, northeast Iowa. The American Midland Naturalist 100:307-319.
- Curtis, J. T. 1959. The vegetation of Wisconsin: An ordination of plant communities. Reprinted in 1987. University of Wisconsin Press, Madison. 657 pp.
- Faber-Langendoen, D., editor. 2001. Plant communities of the Midwest: Classification in an ecological context. Association for Biodiversity Information, Arlington, VA. 61 pp. plus appendix (705 pp.).
- INAI [Iowa Natural Areas Inventory]. 2017. Vegetation classification of Iowa. Iowa Natural Areas Inventory, Iowa Department of Natural Resources, Des Moines.
- MNNHP [Minnesota Natural Heritage Program]. 1993. Minnesota''s native vegetation: A key to natural communities. Version 1.5. Minnesota Department of Natural Resources, Natural Heritage Program, St. Paul, MN. 110 pp.
- McIntosh, R. 1950. Pine stands in southwestern Wisconsin. Wisconsin Academy of Arts and Letters 40:243-257.
- Midwestern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Minneapolis, MN.
- Minnesota DNR [Minnesota Department of Natural Resources]. 2003-2005a. Field guide to the native plant communities of Minnesota. Three volumes: The Laurentian Mixed Forest Province (2003), The Eastern Broadleaf Forest Province (2005c), The Prairie Parkland and Tallgrass Aspen Parklands provinces (2005b). Ecological Land Classification Program, Minnesota County Biological Survey, and Natural Heritage and Nongame Research Program. Minnesota Department of Natural Resources, St. Paul.
- WDNR [Wisconsin Department of Natural Resources]. 2015. The ecological landscapes of Wisconsin: An assessment of ecological resources and a guide to planning sustainable management. PUB-SS-1131 2015. Wisconsin Department of Natural Resources, Madison. [http://dnr.wi.gov/topic/landscapes/Book.html]