Print Report

CEGL001332 Coleogyne ramosissima Shrubland

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Blackbrush Shrubland

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This broadly defined, common shrubland association occurs in the Colorado Plateau, southern Great Basin, Mojave Desert, and Sierra Nevada foothills. The vegetation is characterized by an open to moderately dense short-shrub layer that is clearly dominated by the evergreen microphyllous shrub Coleogyne ramosissima, sometimes in nearly pure stands with a typically sparse herbaceous layer. Shrub cover is usually around 20%, although it can range as low as 5% or as high as 50%. Because it is a wide-ranging type, a variety of other shrub and dwarf-shrub species may be present with low cover, including Acamptopappus sphaerocephalus, Ambrosia dumosa, Artemisia bigelovii, Artemisia filifolia, Atriplex canescens, Atriplex confertifolia, Chrysothamnus viscidiflorus, Ephedra funerea, Ephedra cutleri, Ephedra nevadensis, Ephedra torreyana, Ephedra viridis, Ericameria linearifolia, Ericameria teretifolia, Eriogonum corymbosum, Gutierrezia sarothrae, Gutierrezia microcephala, Krascheninnikovia lanata, Lycium spp., Menodora spinescens, Opuntia spp., Psorothamnus fremontii, and Yucca baccata. Occasional Juniperus spp., Pinus edulis, or Pinus monophylla trees are present in some stands. The herbaceous layer generally includes only sparse cover of graminoids and forbs, except during wet years when cover of annuals may be high. Cover of the introduced annual Bromus spp. may be high in disturbed stands, but in general, the substrate does not support the growth of more than a trace of grasses. Stands are found on a variety of landforms. Sites are on level to steep slopes, rarely exceeding 45% and include all aspects. In southern Nevada, stands occur on lower foothills and upper bajadas, often with cooler northern and eastern aspects. In the Colorado Plateau, stands occupy alluvial fans, benches, canyon bottoms, hills, mesas, plains, plateaus, ridges, rims, sand dunes, sideslopes, terraces, valley bottom fill and colluvial slopes oriented to any aspect. Elevations range from 1065-2133 m (3500-7000 feet). Climate is temperate, continental, with hot summers and cold winters. The unvegetated surface is dominated by bare ground, except on some sandy sites, where biological soil crusts may provide more than 50% cover. Substrates range from barren shales to alluvium and eolian sands to broken limestone. Soils tend to be shallow, calcareous, sandy-textured on eolian sand sites, and clay-textured on shale sites. There is often a caliche subhorizon. Gravel, boulders and rock outcrops are common.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This widespread plant association is defined by the strong dominance of Coleogyne ramosissima without diagnostic codominate shrubs or a developed herbaceous layer. It is similar to the broadly defined ~Coleogyne ramosissima / Pleuraphis jamesii Shrubland (CEGL001334)$$ and is separated from it by the lack of a Pleuraphis jamesii-dominated herbaceous layer. In cases where the grass element is missing or is greatly reduced because of grazing and not because of edaphic factors, there is potential for misclassification of stands.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: This common association is characterized by an open to moderately dense short-shrub layer that is clearly dominated by the evergreen microphyllous shrub Coleogyne ramosissima. Shrub cover is usually around 20%, although it can range as low as 5% or as high as 50%. It can occur in almost pure stands. Other shrub and dwarf-shrub species may be present with low cover, including Ambrosia dumosa, Artemisia filifolia, Atriplex canescens, Atriplex confertifolia, Chrysothamnus viscidiflorus, Ephedra funerea, Ephedra torreyana, Ephedra nevadensis, Ephedra viridis, Ericameria linearifolia, Ericameria teretifolia, Gutierrezia sarothrae, Gutierrezia microcephala, Krascheninnikovia lanata, Lycium spp., Menodora spinescens, Opuntia spp., and Yucca baccata. Locally, disturbance may reduce abundance of Coleogyne ramosissima and favor Artemisia filifolia until it nearly codominates. Occasional Juniperus spp., Pinus edulis, or Pinus monophylla trees are present in some stands. The herbaceous layer generally includes only sparse cover of graminoids and forbs, except during wet years when cover of annuals may be high. Cover of the introduced annual Bromus spp. may be high in disturbed stands, but in general, the substrate does not support the growth of more than a trace of grasses. Forbs may include species of Chaenactis, Cryptantha, Phacelia, Plantago, and Streptanthella. Scattered Juniperus osteosperma may also be present.

Dynamics:  Coleogyne ramosissima is very sensitive to fire because it does not resprout after burning, and seeds in the seed bank are short-lived and destroyed by fire (Bowns and West 1976, Wright 1980). Many stands are too sparse to burn, but if burned, Coleogyne ramosissima is slow to recolonize, often requiring more than 60 years to re-establish (Wright 1980). Burned-over areas convert to Gutierrezia microcephala- or Artemisia tridentata-dominated shrublands (Bowns and West 1976). Invasion of the introduced annual Bromus spp. creates a fire hazard and increases fire frequency (Warren et al. 1982).

Environmental Description:  This broadly defined shrubland association occurs in the Colorado Plateau, southern Great Basin, Mojave Desert, and Sierra Nevada foothills in areas with hot summers and cold winters. Elevations range from 1065-2133 m (3500-7000 feet). Sites are frequently on level to moderate slopes rarely exceeding 20% but have also been found on steep slopes (>45%). In Nevada, stands occur on lower foothills and upper bajadas, often with cooler northern and eastern aspects. In the Colorado Plateau, stands occupy plateaus, ridges, dunes, alluvial fans, canyon rims, bench ledges valleys, plains, and colluvial slopes with any aspect. The upper elevation ecotones are generally narrow, and Coleogyne ramosissima may mix with Artemisia spp. or Pinus edulis and Juniperus osteosperma. The lower elevation ecotones tend to be broader, and Coleogyne ramosissima may mix with grasslands or with shrublands dominated by Larrea tridentata, Ambrosia dumosa, Atriplex spp., or Grayia spinosa (Beatley 1976). Stands described from the Colorado Plateau often occur in a mosaic with pinyon-juniper woodlands (Warren et al. 1982). The unvegetated surface is dominated by bare ground, except on some sandy sites, where biological soil crusts may provide more than 50% cover. Substrates range from barren shales to alluvium and eolian sands to broken limestone. Soils tend to be shallow, calcareous, sandy-textured on eolian sand sites, and clay soils on shale sites. There is often with a caliche subhorizon. Gravel, boulders and rock outcrops are common.

Geographic Range: This widespread and common blackbrush shrubland association occurs in the Colorado Plateau, southern Great Basin, Mojave Desert, and Sierra Nevada foothills. It ranges from northern Arizona to Utah west to California and Nevada.

Nations: US

States/Provinces:  AZ, CA, NV, UT




Confidence Level: High

Confidence Level Comments: No Data Available

Grank: G4G5

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = 152.1311 Coleogyne ramosissima - Ephedra viridis - Yucca baccata (Warren et al. 1982)
= Coleogyne ramosissima - Ephedra nevadensis Association (Peterson 1984a) [Occurs within Peterson''s (1984) Mixed Shrub Zone. Eriogonum fasciculatum was present in some of the stands.]
= Coleogyne ramosissima - Ephedra nevadensis Shrubland (Ostler et al. 2000)
= Coleogyne ramosissima (Sawyer et al. 2009) [33.020.01]
= Coleogyne ramosissima (Evens et al. 2014)
? Coleogyne ramosissima Association (Annable 1985) [Occurs within Annable''s (1985) Mixed Shrub Zone.]
< Coleogyne ramosissima Communities (Beatley 1976)
= Blackbrush Plant Community (Armstrong 1969)
= Blackbrush Type (Schultz et al. 1987)
< Desert Shrub (BIA 1979)
= Mohavean Shrub (Coleogyne) Formation (Wells 1960)

Concept Author(s): K.A. Schulz

Author of Description: K.A. Schulz and J. Coles

Acknowledgements: No Data Available

Version Date: 07-14-16

  • Annable, C. R. 1985. Vegetation and flora of the Funeral Mountains, Death Valley National Monument. California-Nevada Cooperative National Park Resources Studies Unit, National Park Service/University of Nevada Contribution 016/07. Las Vegas, NV. 188 pp.
  • Armstrong, J. D. 1969. Vegetation of the Virgin Mountains, Clark County, Nevada. Unpublished thesis, University of Nevada, Las Vegas. 104 pp.
  • BIA [Bureau of Indian Affairs]. 1979. The secretarial land use plan for the addition to the Havasupai Indian Reservation. Unpublished draft Environmental Statement INT DES 79-42. Prepared by USDI Bureau of Indian Affairs, Phoenix Area Office with the assistance of Office of Arid Land Studies, University of Arizona, Tucson.
  • Beatley, J. C. 1976. Vascular plants of the Nevada Test Site and central-southern Nevada: Ecological and geographic distributions. Technical Information Center, Energy Research and Development Administration. TID-26881. Prepared for Division of Biomedical and Environmental Research. 297 pp.
  • Bourgeron, P. S., and L. D. Engelking, editors. 1994. A preliminary vegetation classification of the western United States. Unpublished report. The Nature Conservancy, Western Heritage Task Force, Boulder, CO. 175 pp. plus appendix.
  • Bowns, J. E., and N. E. West. 1976. Blackbrush (Coleogyne ramosissima Torr.) on southwestern Utah rangelands. Utah Agricultural Experiment Station Research Report 27. Logan, UT. 27 pp.
  • Bradley, W. G. 1964. The vegetation of the desert game range with special reference to the desert bighorn. Pages 43-67 in: Transcripts of the Desert Bighorn Council. Las Vegas, NV.
  • Callison, J., Jr., J. D. Brotherson, and J. E. Bowns. 1985. The effects of fire on the blackbrush (Coleogyne ramosissima) community of southwestern Utah. Journal of Range Management 38(6):535-538.
  • Clark, D., M. Dela Cruz, T. Clark, J. Coles, S. Topp, A. Evenden, A. Wight, G. Wakefield, and J. Von Loh. 2009. Vegetation classification and mapping project report, Capitol Reef National Park. Natural Resource Report NPS/NCPN/NRTR--2009/187. National Park Service, Fort Collins, CO. 882 pp.
  • Cogan, D., M. Reid, K. Schulz, and M. Pucherelli. 2004. Zion National Park, Utah 1999-2003. Vegetation Mapping Project. Technical Memorandum 8260-03-01. Remote Sensing and GIS Group Technical Service Center, Bureau of Reclamation, Denver, CO. Appendix F: Vegetation Association Descriptions for Zion.
  • Coles, J., A. Tendick, G. Manis, A. Wight, G. Wakefield, J. Von Loh, and A. Evenden. 2009a. Vegetation classification and mapping project report, Arches National Park. Natural Resource Technical Report NPS/NCPN/NRTR--2009/253. National Park Service, Fort Collins, CO. 544 pp.
  • Evens, J. M., K. Sikes, D. Hastings, and J. Ratchford. 2014. Vegetation alliance descriptions for Lake Mead National Recreation Area, Death Valley National Park and Mojave National Preserve. Unpublished report submitted to USDI National Park Service, Mojave Desert Network Inventory and Monitoring Program. California Native Plant Society, Sacramento, CA.
  • Kearsley, M. J. C., K. Green, M. Tukman, M. Reid, M. Hall, T. J. Ayers, and K. Christie. 2015. Grand Canyon National Park-Grand Canyon / Parashant National Monument vegetation classification and mapping project. Natural Resource Report NPS/GRCA/NRR--2015/913. National Park Service, Fort Collins, CO. 75 pp. plus appendices.
  • Ostler, W. K., D. J. Hansen, D. C. Anderson, and D. B. Hall. 2000. Classification of vegetation on the Nevada Test Site. DOE/NV/11718-477. U.S. Department of Energy, Bechtel Nevada Ecological Services, Las Vegas, NV. 102 pp.
  • Peterson, E. B. 2008. International Vegetation Classification alliances and associations occurring in Nevada with proposed additions. Nevada Natural Heritage Program, Carson City, NV. 348 pp.
  • Peterson, P. M. 1984a. Flora and physiognomy of the Cottonwood Mountains, Death Valley National Monument, California. University of Nevada Cooperative National Park Resources Studies Unit Report CPSU/UNLV 022/06. Las Vegas, NV.
  • Reid, M. S., and M. E. Hall. 2010. Vegetation classification of Grand Canyon National Park. Draft report submitted to National Park Service, Fort Collins, CO.
  • Romme, W. H., K. D. Heil, J. M. Porter, and R. Fleming. 1993. Plant communities of Capitol Reef National Park, Utah. USDI National Park Service, Technical Report NPS/NAUCARE/NRTER-93/02. Cooperative Park Studies Unit, Northern Arizona University. 37 pp.
  • Sawyer, J. O., T. Keeler-Wolf, and J. Evens. 2009. A manual of California vegetation. Second edition. California Native Plant Society, Sacramento CA. 1300 pp.
  • Schultz, L. M., E. E. Neely, and J. S. Tuhy. 1987. Flora of the Orange Cliffs of Utah. Great Basin Naturalist 47:287-298.
  • Shields, L. M., W. H. Rickard, and F. Drouet. 1959. A botanical study of nuclear effects at the Nevada Test Site. 1958 Annual Report, New Mexico Highlands University, to U.S. Atomic Energy Commission.
  • Spolsky, A. M. 1979. An overview of the plant communities of Anza Borrego Desert State Park. Unpublished report. State of California, The Resource Agency, Department of Parks and Recreation , Anza-Borrego Desert State Park, Borrego Springs, CA.
  • Tendick, A., J. Coles, K. Decker, M. Hall, J. Von Loh, T. Belote, A. Wight, G. Wakefield, and A. Evenden. 2012. Vegetation classification and mapping project report, Canyonlands National Park. Natural Resource Technical Report NPS/NCPN/NRTR--2012/577. National Park Service, Fort Collins, CO.
  • Tendick, A., J. Spence, M. Reid, K. Shulz, G. Kittel, K. Green, A. Wight, and G. Wakefield. 2017. Vegetation classification and mapping project report, Glen Canyon National Recreation Area and Rainbow Bridge National Monument. Natural Resource Report NPS/NCPN/NRR—2017/1500. National Park Service, Fort Collins, CO. 1464 pp.
  • Warren, P. L., K. L. Reichhardt, D. A. Mouat, B. T. Brown, and R. R. Johnson. 1982. Vegetation of Grand Canyon National Park. Cooperative National Park Resources Studies Unit Technical Report 9. Tucson, AZ. 140 pp.
  • Wells, P. V. 1960. Physiognomic intergradation of vegetation on the Pine Valley Mountains in southwestern Utah. Ecology 41:553-556.
  • West, N. E. 1983d. Colorado Plateau-Mohavian blackbrush semi-desert. Pages 399-412 in: N. E. West, editor. Temperate deserts and semi-deserts. Ecosystems of the world, Volume 5. Elsevier Publishing Company, Amsterdam.
  • Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.
  • Wright, H. A. 1980. The role and use of fire in the semi-desert grass-shrub type. General Technical Report INT-85. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 23 pp.