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CEGL008435 (Hydrangea arborescens) / Heuchera villosa - Asplenium trichomanes - Thalictrum clavatum / Conocephalum salebrosum Cliff Shrubland
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: (Wild Hydrangea) / Hairy Alumroot - Maidenhair Spleenwort - Mountain Meadowrue / Snakeskin Liverwort Cliff Shrubland
Colloquial Name: Southern Appalachian Moist Siltstone Cliff
Hierarchy Level: Association
Type Concept: This type occurs on moist, shaded, north-facing cliffs of thin-bedded sedimentary rocks (siltstones and shales) in the Ridge and Valley and the adjacent western portion of the Blue Ridge in southwestern Virginia and Tennessee. It has sparse to moderately dense cover of vascular plants and also has substantial cover of nonvascular plants. Vascular plants root in the horizontal crevices. Most abundant vascular species in the Tennessee occurrences are Heuchera villosa var. villosa, Asplenium trichomanes ssp. trichomanes, and Thalictrum clavatum. Other vascular species present include Hydrangea arborescens, Rhododendron maximum, Laportea canadensis, Trautvetteria caroliniensis var. caroliniensis, Dryopteris intermedia, Dryopteris marginalis, Polypodium appalachianum, Asplenium montanum, and (rarely) Phegopteris connectilis and Saxifraga caroliniana. Conocephalum conicum is a common and conspicuous nonvascular plant.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: This community is arguably also closely related to ~Appalachian Acidic Cliff Alliance (A2071)$$. It seems better placed here, because of the overall floristic composition, the subcalcareous sedimentary rock substrate, the relatively continuously moist conditions, and the high cover of nonvascular plants. The description and concept of the type may require substantial revision when additional information is available. An example of this community has been documented from the Dickey Branch Preserve (The Nature Conservancy), Johnson County, Tennessee (Plot CHER.4).
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: In Tennessee, this community has sparse to moderately dense cover of vascular plants and also has substantial cover of nonvascular plants. Vascular plants root in the horizontal crevices. Most abundant vascular species are Heuchera villosa var. villosa, Asplenium trichomanes ssp. trichomanes, and Thalictrum clavatum. Other vascular species present include Hydrangea arborescens, Rhododendron maximum, Laportea canadensis, Trautvetteria caroliniensis var. caroliniensis, Dryopteris intermedia, Dryopteris marginalis, Polypodium appalachianum, Asplenium montanum, and (rarely) Phegopteris connectilis and Saxifraga caroliniana. Conocephalum conicum is a common and conspicuous nonvascular plant.
The three Virginia occurrences have similar vegetation, but the full floristic composition and patterns of variation over this cliff will probably never be known due to the extreme inaccessibility of the habitat and the danger involved with inventory. At all three sites, Hydrangea arborescens, Physocarpus opulifolius var. opulifolius, Heuchera villosa var. villosa, Saxifraga caroliniana, Solidago arguta, Campanula divaricata, Dryopteris marginalis, and Sedum ternatum are important species. Other species that could be observed on one or two of the cliffs include Cystopteris bulbifera, Packera obovata, Solidago ulmifolia var. ulmifolia, Helianthus divaricatus, Carex pensylvanica, and Arabis laevigata var. laevigata. Stunted, shrub-sized trees of Acer saccharum, Fraxinus americana, and Carya glabra are also scattered in crevices, especially near the tops of the cliffs. Vegetation ranges from sparse on vertical faces to rather dense on shelves and sloping faces where veneers of soil have accumulated.
The three Virginia occurrences have similar vegetation, but the full floristic composition and patterns of variation over this cliff will probably never be known due to the extreme inaccessibility of the habitat and the danger involved with inventory. At all three sites, Hydrangea arborescens, Physocarpus opulifolius var. opulifolius, Heuchera villosa var. villosa, Saxifraga caroliniana, Solidago arguta, Campanula divaricata, Dryopteris marginalis, and Sedum ternatum are important species. Other species that could be observed on one or two of the cliffs include Cystopteris bulbifera, Packera obovata, Solidago ulmifolia var. ulmifolia, Helianthus divaricatus, Carex pensylvanica, and Arabis laevigata var. laevigata. Stunted, shrub-sized trees of Acer saccharum, Fraxinus americana, and Carya glabra are also scattered in crevices, especially near the tops of the cliffs. Vegetation ranges from sparse on vertical faces to rather dense on shelves and sloping faces where veneers of soil have accumulated.
Dynamics: This community is generally stable. Periodic rock fall occurs.
Environmental Description: This community is known from north-facing, seepy cliffs over siltstones and shales of intermediate composition, at 762-915 m (2500-3000 feet) elevation, in the Ridge and Valley and western Blue Ridge provinces. In the southwestern Virginia Ridge and Valley, three calcareous siltstone cliffs of the Juniata Formation support similar vegetation. The Virginia habitats are massive red siltstone faces ranging from about 40 to 200 m high, with occasional small ledges, cracks, and ephemeral seepage only in local areas. Aspect ranges from northeast to southeast. Elevation lies at about 1130 to 1220 m (3700-4000 feet).
Geographic Range: This type is known from a small area of Ridge and Valley and adjacent Blue Ridge (with sedimentary inclusions) in Virginia and Tennessee. Its potential occurrence is more widespread.
Nations: US
States/Provinces: AL?, GA?, TN, VA
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.686196
Confidence Level: Low - Poorly Documented
Confidence Level Comments: No Data Available
Grank: G2
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 6 Open Rock Vegetation Class | C06 | 6 |
Subclass | 6.B Temperate & Boreal Open Rock Vegetation Subclass | S04 | 6.B |
Formation | 6.B.1 Temperate & Boreal Cliff, Scree & Other Rock Vegetation Formation | F034 | 6.B.1 |
Division | 6.B.1.Na Eastern North American Temperate Cliff, Scree & Rock Vegetation Division | D051 | 6.B.1.Na |
Macrogroup | 6.B.1.Na.1 Shrubby Fivefingers - Rock Polypody / Cup Lichen species Eastern North American Cliff & Rock Vegetation Macrogroup | M111 | 6.B.1.Na.1 |
Group | 6.B.1.Na.1.c Appalachian Cliff & Rock Vegetation Group | G840 | 6.B.1.Na.1.c |
Alliance | A2072 Wild Hydrangea / Hairy Alumroot Appalachian Circumneutral Cliff Alliance | A2072 | 6.B.1.Na.1.c |
Association | CEGL008435 (Wild Hydrangea) / Hairy Alumroot - Maidenhair Spleenwort - Mountain Meadowrue / Snakeskin Liverwort Cliff Shrubland | CEGL008435 | 6.B.1.Na.1.c |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: No Data Available
- Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
- NatureServe Ecology - Southeastern United States. No date. Unpublished data. NatureServe, Durham, NC.
- Southeastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Durham, NC.
- Szweykowski, J., K. Buczkowska, and I. J. Odrzykoski. 2005. Conocephalum salebrosum (Marchantiopsida, Conocephalaceae) - a new Holarctic liverwort species. Plant Systematics and Evolution 253:133-158.