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CEGL002102 Quercus phellos - (Quercus lyrata) / Carex spp. - Leersia spp. Pond Forest
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: Willow Oak - (Overcup Oak) / Sedge species - Cutgrass species Pond Forest
Colloquial Name: Willow Oak Bottomland Flatwoods Forest
Hierarchy Level: Association
Type Concept: This willow oak flatwoods forest occurs in bottomland environments of major rivers in the south-central United States. This type most often occurs on first bottoms and terrace flats with poorly drained, heavy silty clay soils that are flooded with shallow water each winter and spring. The relatively high clay content in these soils produces a perched water table that contributes to prolonged wet conditions and equally severe summer dry periods when soils become hard and develop excessive cracks. This restricted soil permeability is characteristic of flatwoods, which contain species typical of drier upland sites on slight rises and shallow ephemeral seasonal ponds in depressions. The perched water table situation can also occur in distinctly upland situations as well as in floodplains. Dominant trees in this community include Quercus phellos, Celtis laevigata, Fraxinus pennsylvanica, and Ulmus americana. Quercus lyrata and Quercus texana are also frequently encountered; Quercus stellata is an occasional codominant. Unfavorable soil conditions often cause the trees to be stunted. The understory and shrub layers are poorly developed (less than 50% cover) consisting primarily of Acer rubrum, Ilex decidua, and Forestiera acuminata. The herbaceous stratum is dominated by Carex spp., Leersia spp., and Cinna arundinacea. Herbaceous density is generally low but fluctuates with canopy openness. Elevated rises exhibit plants common of dry and dry-mesic soils such as Gillenia stipulata and Pycnanthemum tenuifolium, while wet and wet-mesic soils found in shallow ephemeral or seasonal ponds in depressions are dominated by Carex spp. Symphyotrichum lanceolatum is often encountered in flatwoods, as are Toxicodendron radicans and Campsis radicans.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: Large, mature examples of this are reported from Kentucky (M. Evans pers. comm. 2003). A very homogenous and uniform stand with canopy strongly dominated by the two co- dominants, Quercus phellos and Quercus stellata, occurs on a primary terrace of a medium size stream in the Shawnee Hills of Kentucky. It contains sparse mid and understory woody layers, and the herbaceous layer is dense and strongly dominated by sedges (M. Hines pers. comm. 2019). Elsewhere, this community is most prevalent in the poorly drained bottomlands of the Mississippi River and its major tributaries. It can be found in extreme southern Illinois and southeastern Missouri, from where it was initially described; however, in Illinois, Quercus phellos is reported only from Massac County (Mohlenbrock and Voigt 1959), and in Missouri this type could be lumped with ~Quercus palustris - (Quercus stellata) - Quercus pagoda / Isoetes spp. Wet Forest (CEGL002101)$$ (M. Leahy pers. comm. 1999). Further field research is needed to document this community outside the Mississippi River Alluvial Plain. This type or related vegetation has been reported from and attributed to the Interior Low Plateau of Kentucky (J. Campbell pers. comm. 1999), as well as Louisiana (L. Smith pers. comm. 1999), and southern Arkansas (D. Zollner pers. comm. 1999). More information is needed on these occurrences. Its proposed occurrence in Mississippi and Tennessee is speculative.
This type may need to be split as more information becomes available. Similar Quercus phellos flatwoods occur in southern Kentucky''s karst region (Pennyroyal Plateau) and are wetter than other Kentucky flatwoods types (J. Campbell pers. comm. 1999). Given that the alliance concept includes communities primarily found in upland depressions, the placement of this association here is problematic and needs to be assessed. Quercus phellos is the indicator species and should be the dominant contributor to the forest canopy. Another unique diagnostic attribute of this community is the presence of dry and dry-mesic herbaceous species on slight rises and wet or wet-mesic herbaceous plants in shallow ephemeral pools.
This type may need to be split as more information becomes available. Similar Quercus phellos flatwoods occur in southern Kentucky''s karst region (Pennyroyal Plateau) and are wetter than other Kentucky flatwoods types (J. Campbell pers. comm. 1999). Given that the alliance concept includes communities primarily found in upland depressions, the placement of this association here is problematic and needs to be assessed. Quercus phellos is the indicator species and should be the dominant contributor to the forest canopy. Another unique diagnostic attribute of this community is the presence of dry and dry-mesic herbaceous species on slight rises and wet or wet-mesic herbaceous plants in shallow ephemeral pools.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: Structural variation in this community is regulated by the duration and depth of flooding, moisture availability, and soil physical properties. Understory, shrub, herbaceous, and vine strata can be dense when moisture conditions are ideal and canopy closure is incomplete (patchy). Quite often, however, the vegetation in this element shows outward signs (wilting, poor growth) of drought conditions during the hot summer months. Many flatwoods had savanna vegetation in presettlement times (White and Madany 1978). Trees found in this community today often exhibit tall, straight trunks with full, symmetrical crowns. Willow oak (and overcup oak to a lesser extent) has abundant, small, stout, spur-like live and dead lower branches which are similar to the "pins" which decorate the trunks of pin oaks (Quercus palustris). These trees are most often open grown, creating an incomplete canopy (80-90% cover) (Nelson 1985).
Floristics: This community is dominated by broad-leaved deciduous trees and has sparse to well-developed understory, shrub, and herbaceous strata. Dominant trees include Quercus phellos, Celtis laevigata, Fraxinus pennsylvanica, and Ulmus americana. In addition, Quercus lyrata and Quercus texana also frequently occur. Quercus stellata is an occasional codominant. Trees in this forest are usually of medium height (20-30 m) (Nelson 1985), and unfavorable soil conditions often cause them to be stunted. The understory and shrub layers are poorly developed (less than 50% cover) consisting primarily of Acer rubrum, Ilex decidua, and Forestiera acuminata. The herbaceous stratum is dominated by Carex spp., Leersia oryzoides, and Cinna arundinacea. Herbaceous density is generally low but fluctuates with canopy openness. Elevated rises exhibit plants common of dry and dry-mesic soils such as Gillenia stipulata (= Porteranthus stipulatus) and Pycnanthemum tenuifolium, while wet and wet-mesic soils found in shallow ephemeral or seasonal ponds in depressions are dominated by sedges. Symphyotrichum lanceolatum (= Aster lanceolatus) is a ubiquitous species often encountered in flatwoods, as are the vines Toxicodendron radicans and Campsis radicans (Voigt and Mohlenbrock 1964, TNC 1995a).
Considerable variability in species composition and physiognomy is encountered within and among occurrences of this natural community throughout its range. This variability is a result of hydrologic influences (duration, depth, and timing of flooding), soil physical properties (the presence of clay or a fragipan), and the amount of light penetrating the tree canopy. Quercus phellos (Willow oak) is diagnostic and the dominant canopy species in all occurrences of this community.
Considerable variability in species composition and physiognomy is encountered within and among occurrences of this natural community throughout its range. This variability is a result of hydrologic influences (duration, depth, and timing of flooding), soil physical properties (the presence of clay or a fragipan), and the amount of light penetrating the tree canopy. Quercus phellos (Willow oak) is diagnostic and the dominant canopy species in all occurrences of this community.
Dynamics: Natural disturbance includes damage caused by flooding, wind, lightning, drought, and fire. This community is seasonally wet due to the presence of a shallow, perched water table and then dry in the summer when a hardpan or impermeable clay soils prevent replenishment of soil moisture from capillary action and restrict rooting depth. This community occupies a position which is intermediate in the flooding gradient. Mature trees can be killed by prolonged flooding (Yeager 1949). Although species found in this community suggest a mid-successional stage, unique edaphic conditions probably make it a climax forest. The relatively high clay content (sometimes forming a fragipan) in these soils produces a perched water table that contributes to prolonged wet conditions and equally severe summer dry periods when soils become hard and develop excessive cracks. This restricted soil permeability is characteristic of flatwoods, which contain both species typical of drier upland sites (found on slight rises) and those typical of shallow ephemeral seasonal ponds (which are found in depressions).
Environmental Description: This willow oak forest occurs in bottomland flatwoods environments of major rivers in the south-central United States. This type most often occurs on first bottoms and terrace flats with poorly drained, heavy silty clay soils that are flooded with shallow water each winter and spring. The relatively high clay content (sometimes forming a fragipan) in these soils produces a perched water table that contributes to prolonged wet conditions and equally severe summer dry periods when soils become hard and develop excessive cracks. This restricted soil permeability is characteristic of flatwoods, which contain species typical of drier upland sites on slight rises and shallow ephemeral seasonal ponds in depressions. Some examples (e.g., in Kentucky) are not in bottomlands but are in distinctly upland positions, but otherwise fit the description (M. Evans pers. comm.).
Soils include dominantly level or nearly level soils that formed in water-laid clayey or loamy sediments on the floodplains of the Mississippi and Ohio rivers. Although flatwoods are best developed on soils with massive, brittle fragipans, they are also well represented on relatively impermeable clay soils. In the Midwest, fragipans (hardpan) are usually developed from glacial till of Illinoisan age (White and Madany 1978). This glacial material forms a loamy, brittle subsurface horizon that is low in porosity and organic matter, low or moderate in clay, but high in silt or very fine sand. A fragipan appears cemented and restricts root growth and water flow. Soils high in clay also exhibit low porosity and are slowly to very slowly permeable. Clays also display an affinity for water, which makes it unavailable to plants when soils begin to dry. Clay soils have a high shrink-swell capacity and crack excessively as they dry. During the rainy season, water slowly permeates these soils and ponds in depressions. The soil dries rapidly in the summer or during drought, and herbaceous vegetation wilts in response to this stress. The penetration of tree roots in the subsoil and burrowing by animals is limited by the hardpan (Nelson 1985, White and Madany 1978). Order: Entisol, Mollisol; Subgroup: Typic Fluvaquents, Vertic Haplaquent, Fluvaquentic Halplaquolls; Family: fine-silty, montmorillonitic-mixed, non-acidic, mesic, udic; Series (IL): Belknap, Bonnie, Cape. Bedrock is deeply buried Paleozoic sandstone or stratified Cenozoic marine deposits.
Ponding in shallow depressions encourages the growth of hydrophytic vegetation, while droughty conditions encourage the growth of vegetation typical of drier uplands on slight ridges. Shallow flooding is common, occurring seasonally (winter and spring) for short durations. Tree species in this type may not tolerate deep flooding (Robertson et al. 1984).
Soils include dominantly level or nearly level soils that formed in water-laid clayey or loamy sediments on the floodplains of the Mississippi and Ohio rivers. Although flatwoods are best developed on soils with massive, brittle fragipans, they are also well represented on relatively impermeable clay soils. In the Midwest, fragipans (hardpan) are usually developed from glacial till of Illinoisan age (White and Madany 1978). This glacial material forms a loamy, brittle subsurface horizon that is low in porosity and organic matter, low or moderate in clay, but high in silt or very fine sand. A fragipan appears cemented and restricts root growth and water flow. Soils high in clay also exhibit low porosity and are slowly to very slowly permeable. Clays also display an affinity for water, which makes it unavailable to plants when soils begin to dry. Clay soils have a high shrink-swell capacity and crack excessively as they dry. During the rainy season, water slowly permeates these soils and ponds in depressions. The soil dries rapidly in the summer or during drought, and herbaceous vegetation wilts in response to this stress. The penetration of tree roots in the subsoil and burrowing by animals is limited by the hardpan (Nelson 1985, White and Madany 1978). Order: Entisol, Mollisol; Subgroup: Typic Fluvaquents, Vertic Haplaquent, Fluvaquentic Halplaquolls; Family: fine-silty, montmorillonitic-mixed, non-acidic, mesic, udic; Series (IL): Belknap, Bonnie, Cape. Bedrock is deeply buried Paleozoic sandstone or stratified Cenozoic marine deposits.
Ponding in shallow depressions encourages the growth of hydrophytic vegetation, while droughty conditions encourage the growth of vegetation typical of drier uplands on slight ridges. Shallow flooding is common, occurring seasonally (winter and spring) for short durations. Tree species in this type may not tolerate deep flooding (Robertson et al. 1984).
Geographic Range: This willow oak flatwoods forest occurs in the south-central United States, ranging from extreme southern Illinois and southeast Missouri south to Arkansas, Louisiana, and perhaps elsewhere.
Nations: US
States/Provinces: AL?, AR, IL, KY, MO?, MS, TN
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.686087
Confidence Level: Low - Poorly Documented
Confidence Level Comments: No Data Available
Grank: G3G4Q
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.3 Temperate Flooded & Swamp Forest Formation | F026 | 1.B.3 |
Division | 1.B.3.Na Eastern North American-Great Plains Flooded & Swamp Forest Division | D011 | 1.B.3.Na |
Macrogroup | 1.B.3.Na.2 Pin Oak - Green Ash - Blackgum Swamp Forest Macrogroup | M503 | 1.B.3.Na.2 |
Group | 1.B.3.Na.2.b Overcup Oak - Post Oak - Blackgum Flatwoods & Pond Forest Group | G654 | 1.B.3.Na.2.b |
Alliance | A3429 Willow Oak - Overcup Oak - Water Oak Interior Pond Forest Alliance | A3429 | 1.B.3.Na.2.b |
Association | CEGL002102 Willow Oak - (Overcup Oak) / Sedge species - Cutgrass species Pond Forest | CEGL002102 | 1.B.3.Na.2.b |
Concept Lineage: merged
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: = Quercus phellos - (Quercus lyrata) / Carex spp. - Leersia spp. Forest (Faber-Langendoen 2001)
> Quercus phellos - Quercus palustris shallow floodplain type (Robertson et al. 1984)
< Eastern Broadleaf and Needleleaf Forests: 113: Southern Floodplain Forest (Quercus-Nyssa-Taxodium) (Küchler 1964)
? P1B3c14b. Quercus phellos - Quercus palustris - Quercus lyrata (Foti et al. 1994)
< Sweetgum - Willow Oak: 92 (Eyre 1980)
> Quercus phellos - Quercus palustris shallow floodplain type (Robertson et al. 1984)
< Eastern Broadleaf and Needleleaf Forests: 113: Southern Floodplain Forest (Quercus-Nyssa-Taxodium) (Küchler 1964)
? P1B3c14b. Quercus phellos - Quercus palustris - Quercus lyrata (Foti et al. 1994)
< Sweetgum - Willow Oak: 92 (Eyre 1980)
- Campbell, Julian J. N. Personal communication. Kentucky Field Office, The Nature Conservancy.
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- Southeastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Durham, NC.
- TDNH [Tennessee Division of Natural Heritage]. 2018. Unpublished data. Tennessee Division of Natural Heritage, Nashville, TN.
- TNC [The Nature Conservancy]. 1995a. A classification and description of plant communities in southern Illinois. Report by the Southern Illinois Field Office, Ullin, IL, and the Midwest Regional Office, Minneapolis, MN.
- Voigt, J. W., and R. H. Mohlenbrock. 1964. Plant communities of southern Illinois. Southern Illinois University Press, Carbondale. 202 pp.
- White, J., and M. Madany. 1978. Classification of natural communities in Illinois. Pages 311-405 in: Natural Areas Inventory technical report: Volume I, survey methods and results. Illinois Natural Areas Inventory, Urbana, IL.
- Yeager, J. E. 1949. Effect of permanent flooding in a river-bottom timber area. Bulletin of the Illinois Natural History Survey 25:33-65.
- Zollner, Douglas. Personal communication. Ecologist, The Nature Conservancy, Arkansas Field Office, Little Rock.