Print Report

CEGL006017 Acer saccharum - Quercus muehlenbergii / Cercis canadensis Woodland

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Acer saccharum - Quercus muehlenbergii / Eastern Redbud Woodland

Colloquial Name: Appalachian Sugar Maple - Chinquapin Oak Limestone Woodland

Hierarchy Level:  Association

Type Concept: This calciphilic maple-oak forest is found in the Central Appalachians and adjacent regions of the eastern United States, ranging south and west to the Interior Low Plateau of Tennessee and the Cumberlands of Alabama. Stands occur in thin soils over calcareous substrates, sometimes in association with limestone glades. These are typically closed-canopy, rich, dry to dry-mesic forests; in some stands the canopy may vary from closed to somewhat open, particularly in Pennsylvania at the northern edge of the range. In the Mid-Atlantic region, stands occupying the driest sites are commonly open and somewhat intermediate in physiognomy between forest and woodland. The stands are primarily composed of Acer saccharum or occasionally Acer nigrum, Quercus muehlenbergii, Fraxinus americana, and Ostrya virginiana. Associates include Quercus alba, Tilia americana, Acer nigrum, Ulmus rubra, Celtis occidentalis, Carya ovalis, and Carya ovata. Quercus montana may also be present in some examples. A variable subcanopy and shrub layer contains Cornus florida, Cercis canadensis, Hamamelis virginiana, Rosa carolina, Rhus aromatica, Viburnum prunifolium, Viburnum rafinesqueanum, Viburnum rufidulum, and Zanthoxylum americanum. The sparse to well-developed herb layer may contain Danthonia spicata, Elymus hystrix, Bouteloua curtipendula, Ageratina altissima, Antennaria plantaginifolia, Aquilegia canadensis, Arabis laevigata, Asclepias quadrifolia, Clematis occidentalis (to the north), Houstonia longifolia, Polygonum scandens, Sanicula canadensis, Saxifraga virginiensis, and Packera obovata. Some other herbs recorded in Virginia examples include Agrimonia rostellata, Anemone virginiana var. virginiana, Symphyotrichum patens var. patens, Bromus pubescens, Dichanthelium boscii, Erigeron pulchellus var. pulchellus, Galium circaezans, Sanicula canadensis, Scutellaria elliptica, and Solidago ulmifolia var. ulmifolia. Some stands attributed to this type are mesic forests of steep slopes in the Southern Ridge and Valley which are dominated by Acer saccharum and some combination of Quercus alba and/or Quercus muehlenbergii with Liriodendron tulipifera, Carya spp., and Aesculus flava in either the canopy or subcanopy. The same, or related forests, are reported from limestones of the lower Cumberland Plateau escarpment of Tennessee and possibly Alabama.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: The range of this type, which was initially described from "the High Alleghenies," has gradually extended south to at least the Interior Low Plateau. It may require subdivision. (This type was formerly attributed questionably to the Upper East Gulf Coastal Plain.) Some stands attributed to this type are mesic forests of steep slopes in the Southern Ridge and Valley which are dominated by Acer saccharum and some combination of Quercus alba and/or Quercus muehlenbergii with Liriodendron tulipifera, Carya spp., and Aesculus flava in either the canopy or subcanopy (Andreu and Tukman 1995). The same, or related forests, are reported from limestones of the lower Cumberland Plateau escarpment of Tennessee and possibly Alabama (Bowen et al. 1995). There has been discussion of the merits of subdividing this type, in effect re-splitting former Acer saccharum - Quercus (alba, muehlenbergii) / Aesculus flava Forest (CEGL006136) (or an equivalent) out of it again.

In a regional analysis of 1134 plots (Georgia, North Carolina, Tennessee, Virginia) for the Southern Appalachian portion of the Appalachian Trail (Fleming and Patterson 2009a), only 14 plots from southwestern Virginia were classified as this type. In a 1300-plot analysis of all Virginia montane upland forest and woodland data (Fleming and Patterson 2009b), 23 plots from across western Virginia are classified as this type. The species that have the highest diagnostic values for the latter group (compared to other calcareous forests) are Muhlenbergia sobolifera, Erigeron pulchellus var. pulchellus, Bromus pubescens, Solidago ulmifolia var. ulmifolia, Brachyelytrum erectum, Ulmus rubra, Packera obovata, Viburnum prunifolium, Parthenocissus quinquefolia, Quercus rubra, Ostrya virginiana, Thalictrum dioicum, Fraxinus americana, and Quercus muehlenbergii.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: The tall, closed-canopy stands are primarily composed of Acer saccharum, Quercus muehlenbergii, Fraxinus americana, and Ostrya virginiana. Associates include Quercus alba, Tilia americana, Acer nigrum, Ulmus rubra, Aesculus flava, Celtis occidentalis, Carya ovalis, Carya cordiformis, and Carya ovata. Quercus montana (= Quercus prinus) may also be present in some examples. Throughout the Central Appalachians, stands occupying the driest sites are commonly open and somewhat intermediate in physiognomy between forest and woodland. A variable subcanopy and shrub layer contains Cornus florida, Cercis canadensis, Carpinus caroliniana, Lindera benzoin, Hamamelis virginiana, Rosa carolina, Rhus aromatica, Viburnum prunifolium, Viburnum rafinesqueanum, Viburnum rufidulum, and Zanthoxylum americanum. The sparse to well-developed herb layer is usually species-rich and may contain Danthonia spicata, Elymus hystrix, Bouteloua curtipendula, Ageratina altissima (= Eupatorium rugosum), Antennaria plantaginifolia, Aquilegia canadensis, Arabis laevigata, Asclepias quadrifolia, Clematis occidentalis (to the north), Houstonia longifolia (= Houstonia tenuifolia), Polygonum scandens, Sanicula canadensis, Saxifraga virginiensis, and Packera obovata (= Senecio obovatus). Some other herbs recorded in Virginia examples include Agrimonia rostellata, Anemone virginiana var. virginiana, Symphyotrichum patens var. patens (= Aster patens var. patens), Bromus pubescens, Dichanthelium boscii, Erigeron pulchellus var. pulchellus, Galium circaezans, Sanicula canadensis, Scutellaria elliptica, and Solidago ulmifolia var. ulmifolia. Some additional taxa that are relatively constant in the broad type are Asplenium platyneuron, Botrychium virginianum, Carex blanda, Actaea racemosa (= Cimicifuga racemosa), Galium triflorum, Hybanthus concolor, Maianthemum racemosum, Polygonatum biflorum, Polymnia canadensis, Ranunculus recurvatus, and Sanguinaria canadensis. In addition, Muhlenbergia sobolifera may dominate the herbaceous stratum of some stands.

Dynamics:  No Data Available

Environmental Description:  This association is typically found on upper sideslopes or summits of limestone, dolomite, or marble ridges with dry soils. Along the New River in West Virginia, it is found in small patches on lower to middle gorge slopes eroded from calcareous geologic formations. These are alkaline forests associated with calcareous soils, often surrounding, or in association with, limestone or dolomite glades. The soils are well-drained, dry, and shallow with outcrops or boulders often present. In the Ridge and Valley and Central Appalachians of Virginia (Fleming 1999), stands of this type are evidently confined to substrates weathered from limestone and dolomite, most frequently occupying submesic to subxeric, southeast- to west-facing slopes at relatively low elevations (mean = 569 m [1867 feet]). These habitats usually appear to be quite dry, rocky, and at least somewhat exposed. The stands are most often situated on middle slopes but range into both lower and upper slope topographic positions. The slope shape is typically convex in at least one direction. Soils are yellow-brown to reddish-brown clay loams and silty clay loams; soil reaction ranges from strongly acidic (pH 5.3) to mildly alkaline (pH 7.4), with mean calcium (Ca) levels of 2474 ppm. In soils weathered from dolomite, magnesium (Mg) levels may exceed 1000 ppm (Fleming 1999). Soil chemistry analyzed from the one WV plot at New River Gorge had high pH (6.8) and high levels of total cation-exchange capacity, calcium (4180 ppm), potassium (121 ppm), and magnesium (424 ppm) compared to most other soils nearby.

Geographic Range: This maple - oak forest is found in the Central Appalachians and adjacent regions of the eastern United States, including the Ridge and Valley and Western Allegheny Plateau regions, ranging from Pennsylvania southward to the Interior Low Plateau of Tennessee and the Ridge and Valley of Virginia.

Nations: US

States/Provinces:  AL?, GA, KY, MD, OH, PA, TN, VA, WV




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: G4?

Greasons: No Data Available


Concept Lineage: The concept, attributes, and range of the Acer saccharum - Quercus (alba, muehlenbergii) / Aesculus flava Forest (former CEGL006136, Tellico Pilot Project) have been (uncomfortably) merged into this association (CEGL006017).

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Acer saccharum - Quercus (alba, muehlenbergii) Forest (Andreu and Tukman 1995)
= Acer saccharum - Quercus muehlenbergii / Cercis canadensis Forest (Fleming and Patterson 2009a)
= Acer saccharum - Quercus muehlenbergii / Houstonia longifolia Forest (Walton et al. 1997)
= Quercus muehlenbergii - Acer (nigrum, saccharum) / Ostrya virginiana / Erigeron pulchellus - Packera obovata Forest (Fleming and Coulling 2001)
> Quercus muehlenbergii / Juniperus virginiana / Hybanthus concolor Association (Rawinski et al. 1996)
= Quercus muhlenbergii - Acer (nigrum, saccharum var. saccharum) / Ostrya virginiana / Senecio obovatus Forest, Type 3.1 (Fleming 1999)
? Yellow oak-sugar maple-red bud forest of calcareous upper slopes and summits (CAP pers. comm. 1998)

Concept Author(s): G.P. Fleming (1999)

Author of Description: L.A. Sneddon, M. Pyne, G.P. Fleming, S.C. Gawler

Acknowledgements: No Data Available

Version Date: 12-15-06

  • Andreu, M. G., and M. L. Tukman. 1995. Forest communities of the Tellico Lake Area, East Tennessee. M.F. project report, Duke University, School of the Environment. Durham, NC. 66 pp. plus appendices.
  • Bartgis, R. L. 1985a. A limestone glade in West Virginia. Bartonia 51:34-36.
  • Bartgis, R. L. 1993. The limestone glades and barrens of West Virginia. Castanea 58:69-89.
  • Bowen, B., M. Pyne, and D. Withers. 1995. An ecological survey of selected tracts in the Tennessee River Gorge: A report to the Tennessee River Gorge Trust. Tennessee Natural Heritage Program, Department of Environment and Conservation, Nashville. 100 pp.
  • CAP [Central Appalachian Forest Working Group]. 1998. Central Appalachian Working group discussions. The Nature Conservancy, Boston, MA.
  • Eastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boston, MA.
  • Fike, J. 1999. Terrestrial and palustrine plant communities of Pennsylvania. Pennsylvania Natural Diversity Inventory. Pennsylvania Department of Conservation and Recreation, Bureau of Forestry, Harrisburg, PA. 86 pp.
  • Fleming, G. P. 1999. Plant communities of limestone, dolomite, and other calcareous substrates in the George Washington and Jefferson national forests, Virginia. Natural Heritage Technical Report 99-4. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. Unpublished report submitted to the USDA Forest Service. 218 pp. plus appendices.
  • Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
  • Fleming, G. P., and K. D. Patterson. 2009a. A vegetation classification for the Appalachian Trail: Virginia south to Georgia. Virginia Department of Conservation and Recreation, Division of Natural Heritage. In-house analysis, March 2009.
  • Fleming, G. P., and K. D. Patterson. 2009b. Classification of selected Virginia montane wetland groups. In-house analysis, December 2009. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
  • Fleming, G. P., and K. Taverna. 2006. Vegetation classification for the National Capitol Region parks, western region. Regional (VA-WVA-MD-DC) analysis prepared for NatureServe and USGS-NPS Vegetation Mapping Program, March 2006. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
  • Fleming, G. P., and P. P. Coulling. 2001. Ecological communities of the George Washington and Jefferson national forests, Virginia. Preliminary classification and description of vegetation types. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. 317 pp.
  • Harrison, J. W. 2011. The natural communities of Maryland: 2011 working list of ecological community groups and community types. Unpublished report. Maryland Department of Natural Resources, Wildlife and Heritage Service, Natural Heritage Program, Annapolis. 33 pp.
  • Harrison, J. W., compiler. 2004. Classification of vegetation communities of Maryland: First iteration. A subset of the International Classification of Ecological Communities: Terrestrial Vegetation of the United States, NatureServe. Maryland Natural Heritage Program, Maryland Department of Natural Resources, Annapolis. 243 pp.
  • ONHD [Ohio Natural Heritage Database]. No date. Vegetation classification of Ohio and unpublished data. Ohio Natural Heritage Database, Division of Natural Areas and Preserves, Ohio Department of Natural Resources, Columbus.
  • Rawinski, T. J., K. N. Hickman, J. Waller-Eling, G. P. Fleming, C. S. Austin, S. D. Helmick, C. Huber, G. Kappesser, F. C. Huber, Jr., T. Bailey, and T. K. Collins. 1996. Plant communities and ecological land units of the Glenwood Ranger District, George Washington and Jefferson national forests, Virginia. Virginia Department of Conservation and Recreation, Division of Natural Heritage. Natural Heritage Technical Report 96-20. Richmond. 65 pp. plus appendices.
  • Vanderhorst, J. P., J. Jeuck, and S. C. Gawler. 2007. Vegetation classification and mapping of New River Gorge National River, West Virginia. Technical Report NPS/NER/NRTR-2007/092. National Park Service, Philadelphia, PA. 396 pp.
  • WVNHP [West Virginia Natural Heritage Program]. No date. Unpublished data. West Virginia Natural Heritage Program, Elkins.
  • Walton, D., N. Putnam, and P. Trianosky. 1997. A classification of the terrestrial plant communities of West Virginia. Second draft. West Virginia Natural Heritage Program, Elkins, WV.