Print Report

CEGL007403 Quercus phellos / Carex (albolutescens, intumescens, joorii) / Climacium americanum Wet Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Willow Oak / (Green-white Sedge, Greater Bladder Sedge, Cypress Swamp Sedge) / American Tree Moss Wet Forest

Colloquial Name: Piedmont Upland Depression Willow Oak Swamp Forest

Hierarchy Level:  Association

Type Concept: This association represents upland depression swamps found in the Piedmont of the Carolinas, Virginia, and the Potomac Valley region of Maryland. It also occurs in limited areas of the adjacent Inner Coastal Plain of the Carolinas. Examples are usually dominated by an almost pure canopy of Quercus phellos, sometimes with codominant Quercus lyrata or Liquidambar styraciflua; scattered individuals of other tree species may be present. Shrubs are generally sparse and not particularly diagnostic of this community. The most characteristic component of the understory are wetland sedges such as Carex albolutescens, Carex intumescens, and Carex joorii, but a number of other species may also be encountered. Sphagnum lescurii, Climacium americanum, and Polytrichum spp. are important moss species. These ponds are important amphibian breeding areas.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Regional classification conducted by VDNH for the NCR and Mid-Atlantic national parks vegetation mapping projects contained a group of 8 Virginia plots and one Maryland plot belonging to this type. The attribution of plots in the Congaree Swamp National Monument was thought to create a contradiction as the Congaree occurrence was thought to be in a depression in a large river floodplain, not in upland depressions, but this is not correct, so there is no problem. It is a mainly Piedmont upland depression pond type, which is also found in the adjacent Inner Coastal Plain of South Carolina (TNC 1998b). This association is also known to occur in Campbell County, Virginia, in the western Piedmont. In contrast to related ponds in central Tennessee, Nyssa biflora is lacking and Cephalanthus occidentalis is uncommon. Carolina Vegetation Survey plot data exist for this association.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: This upland depression swamp forest is usually dominated by an almost pure canopy of Quercus phellos, sometimes with codominant Quercus lyrata, and with Quercus bicolor or Liquidambar styraciflua. In addition, Pinus taeda, Ulmus alata, Quercus stellata, Nyssa biflora, Quercus nigra, Quercus pagoda, Diospyros virginiana, Magnolia virginiana, Quercus michauxii, Ulmus alata, Acer rubrum var. rubrum, and other species may be present. Generally there are patchy or peripheral herbaceous and vine/liana layers of Chasmanthium sessiliflorum, Carex joorii, Rhynchospora glomerata, Mitchella repens, Gelsemium sempervirens, Juncus coriaceus, Smilax tamnoides (= Smilax hispida), Smilax bona-nox, Trachelospermum difforme, Nekemias arborea (= Ampelopsis arborea), Smilax tamnoides, Vitis rotundifolia, Toxicodendron radicans, Bignonia capreolata, and likely others. Shrubs include Ilex decidua, Ilex verticillata, Nyssa sylvatica, Vaccinium fuscatum, Carpinus caroliniana, and abundant Smilax rotundifolia. Common herbs include Carex albolutescens, Carex brevior, Carex caroliniana, Carex crinita, Carex festucacea, Carex intumescens, Carex joorii, Carex louisianica, Carex typhina, Carex squarrosa, Carex pellita (mafic only), Glyceria striata, Scirpus cyperinus, Juncus tenuis, Juncus coriaceus, Chasmanthium laxum, and Trachelospermum difforme. Sphagnum lescurii and Climacium americanum are important moss species. Other Sphagnum spp. and Polytrichum spp. are also found. Exotic plants, especially Lonicera japonica, sometimes become dense in peripheral zones.

Dynamics:  At Congaree Swamp National Monument, South Carolina, this community occurs on about a dozen small depressional features, referred to as "isolated, upland, depressional wetlands." All sites are below three acres in size and are usually oval to circular in shape. In the Piedmont, examples in which the canopy is heavily disturbed sometimes develop successional stands dominated by Liquidambar styraciflua or Acer rubrum.

Environmental Description:  This upland depression swamp forest of the Piedmont and inner Coastal Plain is found in small depressional features with hydrologic regimes controlled by fluctuating groundwater or seasonal pooling of rainwater over impermeable substrates. Sites are usually located on areas underlain by fine-grained mafic rocks, slate, or metasiltstone, and typically have dense clay subsoils that impede drainage. They are typically flooded to a depth up to about 50 cm during the winter and early part of the growing season but draw down by late summer. Below the surficial organic matter, soils are usually gleyed or mottled and have a sticky consistency. Although the parent material is often basic, the soils of these swamps are largely leached of bases and are extremely acidic with moderately low cation levels and base saturation. Occurrences on mafic and acidic substrates appear to be indistinguishable. These are important amphibian breeding areas, but they depend on having intact upland areas as habitat for the adults.

Geographic Range: This vegetation is reported from the Atlantic Coastal Plain of South Carolina and ranges through the Piedmont of the Carolinas and Virginia into the Potomac Valley of Maryland. Throughout its range, this is primarily a Piedmont type.

Nations: US

States/Provinces:  AL, GA?, MD, NC, SC, VA




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: G2G3

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Quercus phellos - Liquidambar styraciflua / Smilax rotundifolia / Carex (albolutescens, festucacea) Woodland (Fleming 2002a)
= Quercus phellos / Carex (intumescens, joorii) - Chasmanthium sessiliflorum / Sphagnum lescurii Forest (Landaal et al. 1998) [Plots 2-57; 2-59; 2-61]
= Quercus phellos / Smilax rotundifolia / Carex (albolutescens, festucacea) Woodland (Fleming and Patterson 2003)
= Upland Depression Swamp Forest (Schafale and Weakley 1990)

Concept Author(s): A.S. Weakley

Author of Description: A.S. Weakley and G.P. Fleming

Acknowledgements: No Data Available

Version Date: 02-16-07

  • ALNHP [Alabama Natural Heritage Program]. 2002. Eufaula National Wildlife Refuge: Natural community and rare plant survey. Alabama Natural Heritage Program, The Nature Conservancy, Montgomery.
  • ALNHP [Alabama Natural Heritage Program]. 2018. Unpublished data on file. Alabama Natural Heritage Program, Auburn University.
  • Fleming, G. P. 2002a. Ecological communities of the Bull Run Mountains, Virginia: Baseline vegetation and floristic data for conservation planning and natural area stewardship. Natural Heritage Technical Report 02-12. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 274 pp. plus appendices.
  • Fleming, G. P. 2002b. Preliminary classification of Piedmont & Inner Coastal Plain vegetation types in Virginia. Natural Heritage Technical Report 02-14. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 29 pp.
  • Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
  • Fleming, G. P., K. Taverna, and P. P. Coulling. 2007b. Vegetation classification for the National Capitol Region parks, eastern region. Regional (VA-MD-DC) analysis prepared for NatureServe and USGS-NPS Vegetation Mapping Program, March 2007. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
  • Fleming, G. P., and K. D. Patterson. 2003. Preliminary vegetation classification for the National Capitol Region parks. Regional (VA-WVA-MD-DC) analysis prepared for NatureServe and USGS-NPS Vegetation Mapping Program, March 2003. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
  • Fleming, G. P., and K. D. Patterson. 2011b. Analysis of Coastal Plain / Outer Piedmont bottomlands and non-alluvial wetlands in Virginia, 400 plots. In-house analysis, January 2011. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
  • Fleming, Gary P. Personal communication. Ecologist, Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA.
  • Harrison, J. W. 2011. The natural communities of Maryland: 2011 working list of ecological community groups and community types. Unpublished report. Maryland Department of Natural Resources, Wildlife and Heritage Service, Natural Heritage Program, Annapolis. 33 pp.
  • Harrison, J. W., compiler. 2004. Classification of vegetation communities of Maryland: First iteration. A subset of the International Classification of Ecological Communities: Terrestrial Vegetation of the United States, NatureServe. Maryland Natural Heritage Program, Maryland Department of Natural Resources, Annapolis. 243 pp.
  • Landaal, S., A. Weakley, and J. Drake. 1998. Classification of the vegetation of Congaree National Park. Report to BRD-NPS Vegetation Mapping Program, National Park Service, The Nature Conservancy, Chapel Hill, NC. 67 pp.
  • Nelson, J. B. 1986. The natural communities of South Carolina: Initial classification and description. South Carolina Wildlife and Marine Resources Department, Division of Wildlife and Freshwater Fisheries, Columbia, SC. 55 pp.
  • Patterson, K. D. 2008a. Vegetation classification and mapping at Appomattox Court House National Historical Park, Virginia. Technical Report NPS/NER/NRTR--2008/125. National Park Service, Philadelphia, PA.
  • Patterson, Karen D. Personal communication. Ecologist, Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA.
  • Peet, R. K., T. R. Wentworth, M. P. Schafale, and A.S. Weakley. No date. Unpublished data of the North Carolina Vegetation Survey. University of North Carolina, Chapel Hill.
  • Schafale, M. P. 2012. Classification of the natural communities of North Carolina, 4th Approximation. North Carolina Department of Environment, Health, and Natural Resources, Division of Parks and Recreation, Natural Heritage Program, Raleigh.
  • Schafale, M. P., and A. S. Weakley. 1990. Classification of the natural communities of North Carolina. Third approximation. North Carolina Department of Environment, Health, and Natural Resources, Division of Parks and Recreation, Natural Heritage Program, Raleigh. 325 pp.
  • Southeastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Durham, NC.