Print Report

CEGL000862 Pinus ponderosa / Muhlenbergia montana Woodland

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Ponderosa Pine / Mountain Muhly Woodland

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This widespread woodland occurs at foothill and lower montane elevations in the southern Rocky Mountains, extending south to the mountains of western Texas, and west to the Mogollon Rim and Colorado Plateau of New Mexico, Arizona and Utah. Elevation ranges from 1890 to 2870 m (6190-9400 feet). Stands occur on bottomlands, elevated plains, cinder cones, piedmont slopes, mesas, foothills, and mountains. Sites include gentle to steep slopes on all aspects, but are more common on southern and western aspects, especially at higher elevations. Substrates are variable but are typically shallow, rocky, coarse-textured soils derived from granitic or cinder parent materials. There is considerable cover of bare soil and exposed bedrock. The vegetation is characterized by an open to moderately dense evergreen, needle-leaved tree canopy to 10-30 m tall that is dominated or codominated by Pinus ponderosa. Associated tree species vary geographically. Pinus edulis, Pinus discolor, and Juniperus spp. may be important in the tree canopy. Pseudotsuga menziesii, Pinus flexilis, and Populus tremuloides may also be present but are considered accidental. Quercus gambelii may be present with low cover (to 5%) in the sparse shrub layer (<10% cover). Other scattered shrubs may include Artemisia tridentata, Ceanothus fendleri, Cercocarpus montanus, Chrysothamnus viscidiflorus, Ericameria nauseosa, Juniperus communis, Purshia tridentata, Rhus trilobata, Ribes cereum, or Yucca baccata. The typically moderately dense herbaceous layer has greater cover than the shrub layer and is dominated by graminoids. Muhlenbergia montana, a warm-season, medium-tall perennial, typically dominates the herbaceous layer and is diagnostic of this association. Common graminoid associates include Aristida spp., Blepharoneuron tricholepis, Bouteloua gracilis, Carex geophila, Carex rossii, Elymus elymoides, Koeleria macrantha, Poa fendleriana, and Schizachyrium scoparium. Festuca arizonica, Muhlenbergia straminea, Muhlenbergia dubia, Muhlenbergia emersleyi, and Hesperostipa spp. are typically absent. Forb cover is typically sparse and highly variable.

Diagnostic Characteristics: Although Muhlenbergia montana is often present to well-represented, it does not have to be present to be called this plant association. Quercus gambelii, if present, is <5% canopy cover. Pseudotsuga menziesii and Populus tremuloides are absent or accidental and then usually in microsites, or if present, may represent an ecotone between this and an adjacent plant association.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This ponderosa pine woodland is a broadly defined plant association. Stuever and Hayden (1997a) suggested the xeric upland and mesic bottomland stands be put into different phases. Fitzhugh et al. (1987) suggested it be divided into regional phases.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: This association is characterized by an open to moderately dense evergreen, needle-leaved tree canopy 10-30 m tall that is dominated or codominated by Pinus ponderosa. Associated tree species vary geographically. Pinus edulis, Pinus discolor, Juniperus monosperma, Juniperus osteosperma, Juniperus deppeana, and Juniperus scopulorum may be important in the tree canopy. Pseudotsuga menziesii, Pinus flexilis, and Populus tremuloides may also be present, but are considered accidental. Quercus gambelii may be present with low cover (to 5%) in the sparse shrub layer (<10% cover). Other scattered shrubs may include Artemisia tridentata, Brickellia californica, Ceanothus fendleri, Cercocarpus montanus, Chrysothamnus viscidiflorus, Ericameria nauseosa, Fallugia paradoxa, Mahonia spp., Purshia tridentata, Quercus grisea, Rhus trilobata, Tetradymia canescens, or Yucca baccata. The typically moderately dense herbaceous layer has greater cover than the shrub layer, and is dominated by graminoids. Muhlenbergia montana, a warm-season, medium-tall perennial, typically dominates the herbaceous layer and is diagnostic of this association. Common graminoid associates include Aristida spp., Blepharoneuron tricholepis, Bouteloua gracilis, Carex geophila, Carex rossii, Elymus elymoides, Koeleria macrantha, Poa fendleriana, and Schizachyrium scoparium. Festuca arizonica, Muhlenbergia straminea (= Muhlenbergia virescens), Muhlenbergia dubia, Muhlenbergia emersleyi, and Hesperostipa spp. are typically absent. Forb cover is typically sparse and may include species such as Antennaria spp., Artemisia ludoviciana, Erigeron spp., Eriogonum racemosum, Chaetopappa ericoides, Lotus wrightii, Oxytropis lambertii, Packera neomexicana, and Penstemon spp.

Dynamics:  Both diagnostic species are tolerant of surface fire. Pinus ponderosa develops thick fire-resistant bark, and Muhlenbergia montana resprouts after burning, although if may take a few years to recover to pre-burn density (Fischer and Bradley 1987, Bradley et al. 1992). This association had frequent fires (every 3-10 years on average) in presettlement times, but fires are less frequent in dry, rocky stands where surface fire is limited by lack of continuous fine fuels (Stuever and Hayden 1997a). Fire-return interval has generally increased because of active fire suppression and historic livestock grazing, which has reduced the fine fuels needed to carry surface fires (Madany and West 1980b, Savage and Swetnam 1990). Absence of fire has led to large accumulations of ground fuel and has likely resulted in denser stands and invasion of less fire-adapted, shade-tolerant species such as Pseudotsuga menziesii. This has likely increased the risk of stand-replacing crown fires.

Improper livestock grazing will favor more grazing-tolerant species, such as Bouteloua gracilis, and over time may eliminate Muhlenbergia montana and convert the stand into ~Pinus ponderosa / Bouteloua gracilis Woodland (CEGL000848)$$.

Environmental Description:  This widespread woodland occurs at foothill and lower montane elevations in the southern Rocky Mountains, extending south to the mountains of western Texas, and west to the Mogollon Rim and Colorado Plateau. Elevation ranges from 1890-2870 m (6190-9400 feet). Stands occur on bottomlands, elevated plains, cinder cones, piedmont slopes, mesas, foothills, and mountains. Sites include gentle to steep slopes on all aspects, but are more common on southern and western aspects, especially at higher elevations. Substrates are variable, but are typically shallow, rocky, coarse-textured soils derived from granitic or cinder parent materials. There is considerable cover of bare soil and exposed bedrock.

Geographic Range: This widespread woodland occurs at foothill and lower montane elevations in the southern Rocky Mountains, extending south to the mountains of western Texas, and west to the Mogollon Rim and Colorado Plateau of New Mexico, Arizona and Utah.

Nations: US

States/Provinces:  AZ, CO, NM, TX, UT




Confidence Level: High

Confidence Level Comments: No Data Available

Grank: G4G5

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Pinus ponderosa - Pseudotsuga menziesii / Muhlenbergia montana Plant Association (Johnston 1987)
= Pinus ponderosa / Muhlenbergia montana (Stuever and Hayden 1997a)
? Pinus ponderosa / Muhlenbergia montana Cover Type (Balice et al. 1997)
= Pinus ponderosa / Muhlenbergia montana Habitat Type (Wasser and Hess 1982)
= Pinus ponderosa / Muhlenbergia montana Habitat Type (Muldavin et al. 1996)
= Pinus ponderosa / Muhlenbergia montana Habitat Type (Alexander et al. 1987)
= Pinus ponderosa / Muhlenbergia montana Habitat Type (Hess and Alexander 1986)
= Pinus ponderosa / Muhlenbergia montana Habitat Type (DeVelice et al. 1986)
= Pinus ponderosa / Muhlenbergia montana Habitat Type (Fitzhugh et al. 1987)
= Pinus ponderosa / Muhlenbergia montana Habitat Type (Hess 1981)
= Pinus ponderosa / Muhlenbergia montana Habitat Type (Youngblood and Mauk 1985)
= Pinus ponderosa / Muhlenbergia montana Plant Association (Baker 1984a)
= Pinus ponderosa / Muhlenbergia montana Plant Association (Larson and Moir 1987)
= Pinus ponderosa / Poa longiligula community type (Hanks et al. 1983)
>< Xeric Foothill Woodland (A3) (Peet 1981)
>< Xeric Montane Woodland (A5) (Peet 1981)

Concept Author(s): M.C. Stuever and J.S. Hayden (1997a)

Author of Description: K.A. Schulz

Acknowledgements: No Data Available

Version Date: 10-24-18

  • Alexander, B. G., Jr., E. L. Fitzhugh, F. Ronco, Jr., and J. A. Ludwig. 1987. A classification of forest habitat types of the northern portion of the Cibola National Forest, NM. General Technical Report RM-143. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 35 pp.
  • Baker, W. L. 1984a. A preliminary classification of the natural vegetation of Colorado. Great Basin Naturalist 44(4):647-676.
  • Balice, R. G., S. G. Ferran, and T. S. Foxx. 1997. Preliminary vegetation and land cover classification for the Los Alamos region. Report LA-UR-97-4627, Los Alamos National Laboratory, Los Alamos, NM.
  • Bourgeron, P. S., and L. D. Engelking, editors. 1994. A preliminary vegetation classification of the western United States. Unpublished report. The Nature Conservancy, Western Heritage Task Force, Boulder, CO. 175 pp. plus appendix.
  • Bradley, A. F., N. V. Noste, and W. C. Fischer. 1992a. Fire ecology of forests and woodlands in Utah. General Technical Report INT-287. USDA Forest Service, Intermountain Research Station, Ogden, UT. 128 pp.
  • CNHP [Colorado Natural Heritage Program]. 2006-2017. Tracked natural plant communities. Colorado Natural Heritage Program, Colorado State University, Fort Collins. [https://cnhp.colostate.edu/ourdata/trackinglist/plant_communities/]
  • DeVelice, R. L. 1983. Forest vegetation of northern New Mexico and southern Colorado. Unpublished dissertation, New Mexico State University, Las Cruces. 191 pp.
  • DeVelice, R. L., J. A. Ludwig, W. H. Moir, and F. Ronco, Jr. 1986. A classification of forest habitat types of northern New Mexico and southern Colorado. General Technical Report RM-131. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 59 pp.
  • DeVelice, R. L., and J. A. Ludwig. 1983a. Climax forest series of northern New Mexico and southern Colorado. Pages 45-53 in: Proceedings of the Workshop on Southwestern Habitat Types, 6-8 April 1983, Albuquerque, NM. USDA Forest Service, Southwest Region, Albuquerque, NM.
  • Diamond, D. D. 1993. Classification of the plant communities of Texas (series level). Unpublished document. Texas Natural Heritage Program, Austin. 25 pp.
  • Fischer, W. C., and A. F. Bradley. 1987. Fire ecology of western Montana forest habitat types. General Technical Report INT-223. USDA Forest Service, Intermountain Research Station, Ogden, UT. 95 pp.
  • Fitzhugh, E. L., W. H. Moir, J. A. Ludwig, and F. Ronco, Jr. 1987. Forest habitat types in the Apache, Gila, and part of the Cibola national forests. General Technical Report RM-145. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 116 pp.
  • Hanks, J. P., E. L. Fitzhugh, and S. R. Hanks. 1983. A habitat type classification system for ponderosa pine forests of northern Arizona. General Technical Report RM-97. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 22 pp.
  • Hansen, M., J. Coles, K. A. Thomas, D. Cogan, M. Reid, J. Von Loh, and K. Schulz. 2004a. USGS-NPS National Vegetation Mapping Program: Walnut Canyon National Monument, Arizona, vegetation classification and distribution. U.S. Geological Survey Technical Report. Southwest Biological Science Center, Flagstaff, AZ. 219 pp.
  • Hansen, M., J. Coles, K. A. Thomas, D. Cogan, M. Reid, J. Von Loh, and K. Schulz. 2004c. USGS-NPS Vegetation Mapping Program: Sunset Crater National Monument, Arizona, vegetation classification and distribution. U.S. Geological Survey Technical Report. Southwest Biological Science Center, Flagstaff, AZ. 188 pp.
  • Hess, K. 1981. Phyto-edaphic study of habitat types of the Arapaho-Roosevelt National Forest, Colorado. Unpublished dissertation, Colorado State University, Fort Collins. 558 pp.
  • Hess, K., and R. R. Alexander. 1986. Forest vegetation of the Arapaho and Roosevelt national forests in northcentral Colorado: A habitat type classification. Research Paper RM-266. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 48 pp.
  • Hibner, C. D. 2009. Special project soil survey of Bandelier National Monument. Natural Resources Conservation Science. In cooperation with the USDI National Park Service and the New Mexico Agricultural Experiment Station. [in review]
  • Johnston, B. C. 1987. Plant associations of Region Two: Potential plant communities of Wyoming, South Dakota, Nebraska, Colorado, and Kansas. R2-ECOL-87-2. USDA Forest Service, Rocky Mountain Region. Lakewood, CO. 429 pp.
  • Larson, M., and W. H. Moir. 1987. Forest and woodland habitat types of northern New Mexico and northern Arizona. Edition 2. USDA Forest Service, Southwestern Region, Albuquerque, NM.
  • Madany, M. H., and N. E. West. 1980b. Fire history of two montane forest areas of Zion National Park. Pages 50-56 in: M. A. Stokes and J. H. Dieterich, technical coordinators. Proceedings of the fire history workshop; 1980 October 20-24; Tucson, AZ. General Technical Report RM-81. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO.
  • Muldavin, E. H., R. L. DeVelice, and F. Ronco, Jr. 1996. A classification of forest habitat types of southern Arizona and portions of the Colorado Plateau. General Technical Report RM-GTR-287. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 130 pp.
  • Muldavin, E., A. Kennedy, C. Jackson, P. Neville, T. Neville, K. Schulz, and M. Reid. 2011b. Vegetation classification and map: Bandelier National Monument. Natural Resource Technical Report NPS/SCPN/NRTR--2011/438. National Park Service, Fort Collins, CO.
  • Muldavin, E., Y. Chauvin, T. Neville, P. Neville, A. Kennedy, H. Hulse, P. Arbetan, K. Schultz, M. Hall, and M. Reid. 2013c. Vegetation classification and map: El Malpais National Park. Natural Resource Technical Report NPS/SCPN/NRTR--2013/803. National Park Service, Fort Collins, CO. [http://nhnm.unm.edu/vlibrary/pubs_archive/nhnm/nonsensitive/R13MUL01NMUS.pdf]
  • Peet, R. K. 1975. Forest vegetation of the east slope of the northern Colorado Front Range. Unpublished dissertation, Cornell University, Ithaca, NY.
  • Peet, R. K. 1981. Forest vegetation of the Colorado Front Range. Vegetatio 45:3-75.
  • Salas, D. E., J. Stevens, K. Schulz, M. Artmann, B. Friesen, S. Blauer, E. W. Schweiger, and A. Valdez. 2010b. Vegetation classification and mapping project report: Great Sand Dunes National Park and Preserve. Natural Resource Report NPS/ROMN/NRR--2010/179. National Park Service, Fort Collins, CO.
  • Salas, D., J. Stevens, and K. Schulz. 2005. USGS-NPS Vegetation Mapping Program: Rocky Mountain National Park, Colorado. Technical Memorandum No. 8260-05-02. USDI Bureau of Reclamation, Denver, CO. 161 pp. plus Appendices A-L (733 pp.).
  • Savage, M., and T. W. Swetnam. 1990. Early 19th-century fire decline following sheep pasturing in a Navajo ponderosa pine forest. Ecology 71(6)2374-2378.
  • Stuever, M. C., and J. S. Hayden. 1997a. Plant associations of Arizona and New Mexico, edition 3. Volume 1: Forests. USDA Forest Service, Southwestern Region. Habitat Typing Guides. 291 pp.
  • Stuever, M. C., and J. S. Hayden. 1997b. Plant associations of Arizona and New Mexico. Volume 2: Woodlands. USDA Forest Service, Southwestern Region, Habitat Typing Guides. 196 pp.
  • TNHS [Texas Natural History Survey]. No date. Unpublished data. Texas Natural History Survey, The Nature Conservancy, San Antonio.
  • Terwilliger, C., K. Hess, and C. Wasser. 1979a. Key to the preliminary habitat types of Region 2. Addendum to initial progress report for habitat type classification. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station. Fort Collins, CO.
  • USFS [U.S. Forest Service]. 1986. Forest and woodland habitat types (plant associations) of southern New Mexico and central Arizona (north of the Mogollon Rim). USDA Forest Service, Southwestern Region, Albuquerque, NM. Second edition, 140 pp. plus insert.
  • USFS [U.S. Forest Service]. 1987a. Forest and woodland habitat types (plant associations) of northern New Mexico and northern Arizona. USDA Forest Service, Southwestern Region, Albuquerque, NM. Second edition, 170 pp. plus insert.
  • USFS [U.S. Forest Service]. 1987b. Forest and woodland habitat types (plant associations) of Arizona south of the Mogollon Rim and southwestern New Mexico. USDA Forest Service, Southwestern Region, Albuquerque, NM. Second edition, 168 pp. plus insert.
  • Wasser, C. H., and K. Hess. 1982. The habitat types of Region II. USDA Forest Service: A synthesis. Final report prepared for USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 140 pp.
  • Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.
  • Youngblood, A. P., and R. L. Mauk. 1985. Coniferous forest habitat types of central and southern Utah. General Technical Report INT-187. USDA Forest Service, Intermountain Research Station, Ogden, UT. 89 pp.