Print Report

CEGL008501 Picea rubens / Betula alleghaniensis / Bazzania trilobata Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Red Spruce / Yellow Birch / Three-lobed Whipwort Forest

Colloquial Name: Red Spruce Forest (Central Appalachian Upland Type)

Hierarchy Level:  Association

Type Concept: This association occurs on both gentle slopes bordering high-elevation valley floors and on more exposed ridge crests and rocky summits in the high Allegheny Mountains of West Virginia and Virginia. Sites range from mesic to submesic and are characterized by cold winter microclimates, frequent fog, and high annual precipitation. Stands are dominated by Picea rubens, which is characteristic, with lower cover of Betula alleghaniensis var. alleghaniensis and Acer rubrum. Other trees that occasionally occur with low cover in the canopy include Tsuga canadensis, Betula lenta, Pinus strobus, and Prunus serotina. The subcanopy is very similar in dominance and composition to the canopy, with the occasional addition of Sorbus americana, Acer pensylvanicum, Amelanchier spp., or Magnolia fraseri. In well-developed stands, young Picea rubens and Betula alleghaniensis var. alleghaniensis are plentiful in understory layers, especially in canopy gaps. Shrub layers are normally somewhat sparse, with low cover by Ilex montana, Kalmia latifolia, Rhododendron maximum, or Vaccinium erythrocarpum. Herb layer composition varies with soil moisture and light regimes but is generally sparse. Dryopteris intermedia, Oxalis montana, Dennstaedtia punctilobula, Maianthemum canadense, and Carex debilis var. rudgei are frequent herbs. The nonvascular stratum is strongly dominated by Bazzania trilobata.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Classification of this unit was revised in 2009 based on 42 plots covering the entire range of this type in West Virginia, as part of a classification and assessment of all red spruce communities in the state. Initial classification of this unit in 2001 was supported by five plot samples from Highland and Rockingham counties, Virginia. Bailey and Ware (1990) and Adams and Stephenson (1991) provide additional quantitative data from Highland County, Virginia, while Stephenson and Clovis (1983) provide data from West Virginia stands. The few stands sampled by Virginia DCR-DNH ecologists show considerable compositional heterogeneity (homogeneity = 0.581), and it is possible that additional plot sampling and analysis would support further subdivision of this unit.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: Picea rubens is the characteristic, dominant canopy tree, with lower cover of Betula alleghaniensis var. alleghaniensis and Acer rubrum. Other trees that occasionally occur with low cover in the canopy include Tsuga canadensis, Betula lenta, Pinus strobus, and Prunus serotina. The subcanopy is very similar in dominance and composition to the canopy, with the occasional addition of Sorbus americana, Acer pensylvanicum, Amelanchier spp., or Magnolia fraseri. In well-developed stands, young Picea rubens and Betula alleghaniensis var. alleghaniensis are plentiful in understory layers, especially in canopy gaps. Shrub layers are normally somewhat sparse, with low cover by Ilex montana, Kalmia latifolia, Rhododendron maximum, or Vaccinium erythrocarpum. Herb layer composition varies with soil moisture and light regimes but is generally sparse. Dryopteris intermedia, Oxalis montana, Dennstaedtia punctilobula, Maianthemum canadense, and Carex debilis var. rudgei are frequent herbs. In Virginia, Lycopodium dendroideum, Lycopodium clavatum, and Oxalis montana are prominent in more mesic situations, and Vaccinium angustifolium and Carex brunnescens ssp. sphaerostachya may be common on exposed, rocky, submesic sites. The nonvascular stratum is strongly dominated by Bazzania trilobata, with lower cover by Hypnum imponens, Dicranum scoparium, Polytrichum pallidisetum, Brotherella recurvans, and Leucobryum glaucum. In West Virginia, two common floristic variants of this community occur. In one variant, shrubs and herbs are almost entirely absent, and the dense spruce canopy shades a carpet of Bazzania liverwort. Another variant is characterized by post-burning increases in Kalmia latifolia, which locally dominates the shrub layer. Species richness of plot-sampled stands ranges from 8 to 31 taxa per 400 m2 (mean = 19). Three Virginia state-rare plants have been documented in this association: Carex arctata, Pyrola elliptica, and Schizachne purpurascens.

Dynamics:  The local invasion of young Picea rubens into northern hardwood understories on middle-slope positions of Allegheny Mountain in Highland County suggests that this association once occupied, or will in the future occupy, a wider variety of habitats in the region (Fleming and Moorhead 1996).

Environmental Description:  This association occurs on both gentle slopes bordering high-elevation valley floors and on more exposed ridge crests and rocky summits. Sites range from mesic to submesic and are characterized by cold winter microclimates, low mean annual temperature, short growing seasons, frequent fog, and high annual precipitation, high relative humidity, and low evapotranspiration rates. Underlying bedrock is sandstone. Soils are extremely acidic (mean pH = 3.4), infertile, frigid silt or sandy loams with thick surficial duff accumulations. Soils are characterized by high organic matter, high cation-exchange capacity, high exchangeable nitrogen, and generally low micronutrient status. This is the typical forest encountered in the heart of red spruce habitat in West Virginia, with widespread distribution in the middle and upper elevations (1070-1400 m [3500-4600 feet]) of the red spruce zone in the state. The local invasion of young Picea rubens into northern hardwood understories on middle-slope positions of Allegheny Mountain in Highland County suggests that this association once occupied, or will in the future occupy, a wider variety of habitats in the region (Fleming and Moorhead 1996).

Geographic Range: This community is nearly restricted to the high Allegheny Mountains of West Virginia and Virginia. In Virginia, scattered stands occur on Allegheny Mountain in northwestern Highland County. There are at least two outliers, at the summit of Sounding Knob (Jack Mountain, Highland County) and on Shenandoah Mountain (Rockingham County) (Stevens 1969), in the adjacent Ridge and Valley region of Virginia. In West Virginia, this community is restricted to five counties (Greenbrier, Pendleton, Pocahontas, Randolph, and Tucker).

Nations: US

States/Provinces:  TN?, VA, WV




Confidence Level: High

Confidence Level Comments: No Data Available

Grank: G2

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Picea rubens / Acer rubrum / Maianthemum canadense - (Lycopodium clavatum, Lycopodium dendroideum) Forest (Fleming and Coulling 2001)
= Picea rubens / Betula alleghaniensis / Bazzania trilobata Forest [Red Spruce - Yellow Birch Forest] (Vanderhorst 2015)
? Picea rubens / Ilex montana / Bazzania trilobata Association (Rawinski et al. 1994)
? Picea rubens / Vaccinium angustifolium - Lycopodium dendroideum Association (Fleming and Moorhead 1996)
< Red Spruce: 32 (Eyre 1980)

Concept Author(s): G.P. Fleming and P. P. Coulling (2001)

Author of Description: G.P. Fleming and E.A. Byers

Acknowledgements: No Data Available

Version Date: 06-01-09

  • Adams, H. S., and S. L. Stephenson. 1991. High elevation coniferous forests in Virginia. Virginia Journal of Science 42:391-399.
  • Allard, H. A., and E. C. Leonard. 1952. The Canaan and the Stony River valleys of West Virginia, their former magnificent spruce forests, their vegetation and floristics today. Castanea 17:1-60.
  • Bailey, C. M., and S. Ware. 1990. Red spruce forests of Highland County, Virginia: Biogeographical considerations. Castanea 55:245-258.
  • Byers, E. A., J. P. Vanderhorst, and B. P. Streets. 2010. Classification and conservation assessment of upland red spruce communities in West Virginia. West Virginia Natural Heritage Program, West Virginia Division of Natural Resources, Elkins.
  • Clarkson, R. B. 1964. Tumult on the mountains: Lumbering in West Virginia - 1770-1920. McClain Printing Co., Parsons, WV. 410 pp.
  • Eastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boston, MA.
  • Eyre, F. H., editor. 1980. Forest cover types of the United States and Canada. Society of American Foresters, Washington, DC. 148 pp.
  • Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
  • Fleming, G. P., and P. P. Coulling. 2001. Ecological communities of the George Washington and Jefferson national forests, Virginia. Preliminary classification and description of vegetation types. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. 317 pp.
  • Fleming, G. P., and W. H. Moorhead, III. 1996. Ecological land units of the Laurel Fork Area, Highland County, Virginia. Natural Heritage Technical Report 96-08. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 114 pp. plus appendices.
  • Murphy, L. S. 1917. The red spruce: Its growth and management. USDA Bulletin 544. Washington, DC. 100 pp.
  • Pielke, R. A. 1981. The distribution of spruce in west-central Virginia before lumbering. Castanea 46:201-216.
  • Rawinski, T. J., G. P. Fleming, and F. V. Judge. 1994. Forest vegetation of the Ramsey''s Draft and Little Laurel Run Research Natural Areas, Virginia: Baseline ecological monitoring and classification. Natural Heritage Technical Report 94-14. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 45 pp. plus appendices.
  • Rollins, A. W. 2005. Analysis of red spruce (Picea rubens) regeneration in Pocahontas, Randolph, and Tucker counties, West Virginia. Master''s thesis, West Virginia University, Morgantown, WV.
  • Schuler, T. M., W. M. Ford, and R. J. Collins. 2002. Successional dynamics and restoration implications of a montane coniferous forest in the Central Appalachians, USA. Natural Areas Journal 22(2):88-98.
  • Stephenson, S. L., and J. F. Clovis. 1983. Spruce forests of the Allegheny Mountains in central West Virginia. Castanea 48:1-12.
  • Stevens, C. E. 1969. A native red spruce stand in Rockingham County. Jeffersonia 3:1-2.
  • Vanderhorst, J. 2015. Wild vegetation of West Virginia: Upland red spruce forests and woodlands. West Virginia Division of Natural Resources, Natural Heritage Program. [http://wvdnr.gov/Wildlife/Factsheets/UplandRedSpruce.shtm]
  • WVNHP [West Virginia Natural Heritage Program]. No date. Unpublished data. West Virginia Natural Heritage Program, Elkins.