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CEGL002443 Pinus resinosa - Pinus strobus / Corylus cornuta / Vaccinium angustifolium Forest
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: Red Pine - White Pine / Beaked Hazelnut / Lowbush Blueberry Forest
Colloquial Name: Subboreal Red Pine - White Pine Dry-Mesic Forest
Hierarchy Level: Association
Type Concept: This upland red pine - white pine community is found in the subboreal regions of the midwestern United States and in central and possibly eastern Canada. This community occurs on Precambrian bedrock, overlaid with medium deep, coarse sand or coarse loam soils. Moisture varies from dry to dry-mesic, and stands are on mid to upper gentle slopes. Stands contain a pine overstory and a poorly developed understory. Pinus resinosa is the dominant overstory species, but can occur mixed with Pinus strobus. Other species include Abies balsamea, Pinus banksiana, Picea mariana, Picea glauca, and Populus tremuloides. Northern hardwoods, such as Acer rubrum and Betula papyrifera, sometimes form a subcanopy. The understory ranges from moderately herb- and shrub-rich to extremely poor. In the tall-shrub strata, the important species are Amelanchier spp. and Corylus cornuta. The short-shrub layer includes Linnaea borealis, Gaultheria procumbens, Vaccinium angustifolium, and Vaccinium myrtilloides, particularly where gaps in the canopy occur. The herb layer is very poorly represented in most parts of this community''s range and includes Aralia nudicaulis, Eurybia macrophylla, Cornus canadensis, Maianthemum canadense, Pteridium aquilinum, and Trientalis borealis. Mosses are abundant, especially Dicranum spp. (including Dicranum polysetum) and Pleurozium schreberi. Diagnostic features include the dominance of Pinus resinosa and the ericaceous understory.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: Stands are dominated by Pinus resinosa, often mixed with Pinus strobus on dry-mesic sites. More mesic white pine and red pine stands are placed into White Pine-Mountain Maple Mesic Forest, ~Pinus strobus / Acer spicatum - Corylus cornuta Forest (CEGL002445)$$. Red pine stands on sand along the Lake Superior shore in Minnesota (FDn32b) and Wisconsin are a separate type [~Pinus banksiana - (Pinus resinosa) - Pinus strobus / Juniperus horizontalis Wooded Grassland (CEGL005125)$$]. Stands in the central Great Lakes region, often on sandy outwash or coarse soils, are separated from this type [~Pinus resinosa - Pinus strobus - (Quercus rubra) / Corylus cornuta Forest (CEGL005399)$$], because of the mix of oak and other northern hardwoods.
A survey of vegetation of the dunes of Lake Michigan in Manistee National Forest in Michigan (Hazlett 1986) found a community very similar to this forest. The described community was closely associated with bogs within the lake dunes. The forest canopy was dominated by Pinus resinosa (75-100% cover), although on the drier ridges inclusions of Quercus velutina and Populus grandidentata were common. Other trees that were found in the canopy included Acer rubrum, Betula papyrifera, Quercus alba, and Quercus rubra. In the understory, species that were found in this community subtype, but not in the main community type, included Gaylussacia baccata, Hamamelis virginiana, and Viburnum nudum var. cassinoides. Many species of herbs were present, but no feathermosses were listed.
A survey of vegetation of the dunes of Lake Michigan in Manistee National Forest in Michigan (Hazlett 1986) found a community very similar to this forest. The described community was closely associated with bogs within the lake dunes. The forest canopy was dominated by Pinus resinosa (75-100% cover), although on the drier ridges inclusions of Quercus velutina and Populus grandidentata were common. Other trees that were found in the canopy included Acer rubrum, Betula papyrifera, Quercus alba, and Quercus rubra. In the understory, species that were found in this community subtype, but not in the main community type, included Gaylussacia baccata, Hamamelis virginiana, and Viburnum nudum var. cassinoides. Many species of herbs were present, but no feathermosses were listed.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: The herb layer is very poorly represented in most parts of this community''s range, contributing only 11% (Ohmann and Ream 1971) of the ground cover in northeastern MN, in contrast with more prominent cover of feather mosses and lichens (35% ground cover) (Grigal and Ohmann 1975).
Floristics: This community is characterized by a pine overstory and a poorly developed understory. Pinus resinosa is present in the canopy at higher cover and basal area than other conifers (typically 80% or more). Pinus strobus can be common in the canopy. Mesic sites tend to include not only Pinus resinosa in the canopy, but also Picea mariana, Picea glauca, and Abies balsamea, whereas on dry sites, Pinus banksiana is found (MNNHP 1993). Beneath the canopy, northern hardwoods, such as Acer rubrum and Betula papyrifera, sometimes form a subcanopy. The understory ranges from moderately herb- and shrub-rich to extremely poor. In the tall-shrub class, the important species are Amelanchier spp. and Corylus cornuta. The short-shrub layer includes Gaultheria procumbens, Linnaea borealis, Vaccinium angustifolium, and Vaccinium myrtilloides, particularly where gaps in the canopy occur (Ohmann and Ream 1971). The herb layer may be moderately closed to sparse, and includes Aralia nudicaulis, Eurybia macrophylla (= Aster macrophyllus), Cornus canadensis, Maianthemum canadense, Pteridium aquilinum, and Trientalis borealis. Mosses can have very high cover and typically include Dicranum spp. (including Dicranum polysetum) and Pleurozium schreberi. The average coverage of feathermoss is greater in the northwest region of Ontario than in the north-central region (Sims et al. 1989). Sims et al. (1989) also found that in the Quetico region of northwestern Ontario, a rich understory of Acer spicatum, Corylus cornuta, Alnus viridis ssp. crispa (= Alnus crispa), Diervilla lonicera, and Aralia nudicaulis was established. Farther north and west, however, the understory was open and dominated by low ericaceous species.
Dynamics: This is an mid-successional community; however, given the appropriate environmental conditions, it can be extremely long-lasting. Older Pinus resinosa trees have the ability to withstand surface fires due to their thick, corky trunks. Pinus resinosa readily colonizes newly exposed, mineral soils, such as those exposed by a fire, provided older trees have survived the fire as a seed source. Given a surface fire regime of perhaps every 20-30 years and crown fires every 100-150 years, this community may sustain itself for hundreds of years (Heinselman 1973). However, Pinus resinosa is not able to germinate under shade and does not appear to be active in gap phase replacement. Therefore, with fire suppression, this community succeeds towards a canopy that includes other conifers or northern hardwoods.
Fire is an important natural disturbance in this community. Pinus resinosa has adaptations that make it well-suited to frequent surface fires and occasional crown fires. Reconstructions of the fire regime (Heinselman 1973, Frelich 1992) in red and white pine stands indicate that a combination of surface fires every 20-30 years and severe crown fires every 100-150 years maintained presettlement Pinus resinosa communities. The thick, corky bark of older Pinus resinosa make them more resistant to fire than any other type of pine. The stems of older Pinus resinosa are likely to have better survival rates than Pinus strobus and Pinus banksiana when no crown damage results (Ahlgren 1974). In addition Pinus resinosa seedling establishment is favored by exposed mineral soils and high sunlight. Therefore, relatively frequent fires makes the continued regeneration of Pinus resinosa possible, as they are only moderately shade-tolerant. Fire suppression over much of the Pinus resinosa range appears to be leading towards the gradual succession from Pinus resinosa forests to forests of mesic hardwoods or Pinus strobus, species that, with a frequent fire regime, are confined to the understory (MNNHP 1993).
Fire is an important natural disturbance in this community. Pinus resinosa has adaptations that make it well-suited to frequent surface fires and occasional crown fires. Reconstructions of the fire regime (Heinselman 1973, Frelich 1992) in red and white pine stands indicate that a combination of surface fires every 20-30 years and severe crown fires every 100-150 years maintained presettlement Pinus resinosa communities. The thick, corky bark of older Pinus resinosa make them more resistant to fire than any other type of pine. The stems of older Pinus resinosa are likely to have better survival rates than Pinus strobus and Pinus banksiana when no crown damage results (Ahlgren 1974). In addition Pinus resinosa seedling establishment is favored by exposed mineral soils and high sunlight. Therefore, relatively frequent fires makes the continued regeneration of Pinus resinosa possible, as they are only moderately shade-tolerant. Fire suppression over much of the Pinus resinosa range appears to be leading towards the gradual succession from Pinus resinosa forests to forests of mesic hardwoods or Pinus strobus, species that, with a frequent fire regime, are confined to the understory (MNNHP 1993).
Environmental Description: This community occurs on Precambrian bedrock, overlaid with medium deep (>60 cm), coarse sand or coarse loam soils. Moisture varies from dry to dry-mesic, and stands are on mid to upper gentle slopes. The midcontinental climate in which this community is found is highly variable, with temperature extremes between -46° and 38°C (average temperature of -11°C in January and 17°C in July), and with anywhere from 58-91 cm annual precipitation. Soil depth ranges from 61-100 cm (Ohmann and Ream 1971). Sims et al. (1989) found that the depth of the organic layer in northwestern Ontario was 6-16 cm for half of the communities sampled and 1-5 cm for the other half. The soil textures are coarse sandy or coarse loamy soils, as well as some fine sands and silts. These soils typically occur over sand dunes or sandy outwash plains. This community occurs predominantly on Precambrian Shield bedrock, mainly on dry to fresh, deep mineral soil. The soil deposition history is both glacio-fluvial and morainal.
Geographic Range: This upland red and white pine community type is found in the hemi-boreal regions of the midwestern United States and in central and possibly eastern Canada, ranging from Minnesota and southwestern Manitoba east to Ontario, and possibly further east.
Nations: CA,US
States/Provinces: MB, MN, ON, QC?
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.684873
Confidence Level: High
Confidence Level Comments: No Data Available
Grank: G3
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.2 Cool Temperate Forest & Woodland Formation | F008 | 1.B.2 |
Division | 1.B.2.Na Eastern North American Forest & Woodland Division | D008 | 1.B.2.Na |
Macrogroup | 1.B.2.Na.6 Eastern White Pine - Jack Pine - Northern Pin Oak Forest & Woodland Macrogroup | M159 | 1.B.2.Na.6 |
Group | 1.B.2.Na.6.c Jack Pine - Red Pine - Black Spruce Subboreal Forest Group | G047 | 1.B.2.Na.6.c |
Alliance | A4130 Red Pine - White Pine Subboreal Forest Alliance | A4130 | 1.B.2.Na.6.c |
Association | CEGL002443 Red Pine - White Pine / Beaked Hazelnut / Lowbush Blueberry Forest | CEGL002443 | 1.B.2.Na.6.c |
Concept Lineage: CEGL002443 & CEGL002520 merged into CEGL002443 (DFL 6-13). Type will be treated as a dune variant of CEGL002444, though it could also be treated as an inland dune variant of CEGL002589, the Great Lakes Dune Pine Forest.
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: = Pinus resinosa / Vaccinium spp. Forest (Faber-Langendoen 2001)
? Pinus resinosa-Populus tremuloides/Diervilla lonicera-Vaccinium spp. Forest (Greenall 1996)
= Northern Dry Forest Red Pine Subtype (Faber-Langendoen 2001)
= Red Pine (Ohmann and Ream 1971)
= Red Pine Forest (MNNHP 1993)
? Pinus resinosa-Populus tremuloides/Diervilla lonicera-Vaccinium spp. Forest (Greenall 1996)
= Northern Dry Forest Red Pine Subtype (Faber-Langendoen 2001)
= Red Pine (Ohmann and Ream 1971)
= Red Pine Forest (MNNHP 1993)
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- CDPNQ [Centre de données sur le patrimoine naturel du Québec]. No date. Unpublished data. Centre de données sur le patrimoine naturel du Québec, Québec.
- Chambers, B. A., B. J. Naylor, J. Nieppola, B. Merchant, and P. Uhlig. 1997. Field guide to forest ecosystems of central Ontario. Southcentral Science Section (SCSS) Field Guide FG-01, Ontario Ministry of Natural Resources, North Bay, Ontario, Canada. 200 pp.
- Faber-Langendoen, D., editor. 2001. Plant communities of the Midwest: Classification in an ecological context. Association for Biodiversity Information, Arlington, VA. 61 pp. plus appendix (705 pp.).
- Farrand, W. R., and D. L. Bell. 1982. Quaternary geology of northern Michigan. University of Michigan, Lansing.
- Frelich, L. E. 1992. The relationship of natural disturbances to white pine stand development. Presented at the White Pine Symposium: History, Ecology, Policy and Management, Duluth, MN. September 16-18, 1992.
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- Heinselman, M. L. 1973. Fire in the virgin forests of the Boundary Waters Canoe Area, Minnesota. Journal of Quaternary Research 3:329-382.
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- Midwestern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Minneapolis, MN.
- Minnesota DNR [Minnesota Department of Natural Resources]. 2003-2005a. Field guide to the native plant communities of Minnesota. Three volumes: The Laurentian Mixed Forest Province (2003), The Eastern Broadleaf Forest Province (2005c), The Prairie Parkland and Tallgrass Aspen Parklands provinces (2005b). Ecological Land Classification Program, Minnesota County Biological Survey, and Natural Heritage and Nongame Research Program. Minnesota Department of Natural Resources, St. Paul.
- ONHIC [Ontario Natural Heritage Information Centre]. 2018. Unpublished data. Ontario Natural Heritage Information Centre, Ministry of Natural Resources, Ontario, Canada.
- Ohmann, L. F., and P. R. Ream. 1971. Wilderness ecology: Virgin plant communities of the Boundary Waters Canoe Area. Research Paper NC-63. USDA Forest Service, North Central Forest Experiment Station, St. Paul, MN. 35 pp.
- Sims, R. A., W. D. Towill, K. A. Baldwin, and G. M. Wickware. 1989. Field guide to the forest ecosystem classification for northwestern Ontario. Ontario Ministry of Natural Resources, Toronto. 191 pp.
- WDNR [Wisconsin Department of Natural Resources]. 2015. The ecological landscapes of Wisconsin: An assessment of ecological resources and a guide to planning sustainable management. PUB-SS-1131 2015. Wisconsin Department of Natural Resources, Madison. [http://dnr.wi.gov/topic/landscapes/Book.html]