Print Report

CEGL006145 Prunus serotina - Sassafras albidum - Amelanchier canadensis - Quercus velutina / Smilax rotundifolia Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Black Cherry - Sassafras - Canadian Serviceberry - Black Oak / Roundleaf Greenbrier Forest

Colloquial Name: Northeastern Maritime Forest

Hierarchy Level:  Association

Type Concept: This maritime forest community of the North Atlantic Coast ecoregion occurs on sheltered backdunes, bluffs, or more interior coastal areas not directly influenced by overwash but affected by salt spray and wind-pruning. Vegetation in these sheltered areas is sometimes referred to as "sunken forest." This name refers to the topographic position of these examples, which are found in large depressions, lower in elevation (by 1-3 m) than the interdunes. These examples are shielded from strong prevailing winds and salt spray, which permits lush growth of broadleaf shrub and vine species. Soils are coarse, well-drained sand subject to considerable shifting during coastal storms, or till and sand deposits of terminal moraines. Physiognomy is variable and ranges from closed-canopy forest to open woodland to dense tall shrubland, and may be more accurately called scrub. Trees found in this community are usually stunted and flat-topped; the canopy may be only 3-7 m tall. Dominant trees vary locally and include Prunus serotina, Sassafras albidum, and Amelanchier canadensis, with admixtures of Celtis occidentalis, Quercus velutina, Pinus rigida, Juniperus virginiana, Acer rubrum, Amelanchier stolonifera, and in southern occurrences Quercus coccinea, Quercus falcata, Liquidambar styraciflua, and Ilex opaca. Additional shrub species may also contribute substantially to the canopy and include Vaccinium corymbosum, Morella pensylvanica, Gaylussacia baccata, Viburnum recognitum, Viburnum dentatum, and Rosa virginiana. A true shrub layer is generally not present. Lianas are common and can be dense in the canopy or the ground layer; species include Parthenocissus quinquefolia, Toxicodendron radicans, Smilax rotundifolia, and Smilax glauca. The understory is generally sparse with tree or vine seedlings plus herbaceous species, including Aralia nudicaulis, Moehringia lateriflora, Maianthemum stellatum, and Maianthemum canadense. Several invasive species can be prevalent in this association, including Lonicera morrowii, Lonicera japonica, Ligustrum vulgare, Berberis vulgaris, and Celastrus orbiculatus.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: ~Prunus serotina / Morella cerifera / Smilax rotundifolia Scrub Forest (CEGL006319)$$ is the southern analog of this association and is differentiated by having species of more southern affinity such as Morella cerifera and Pinus taeda. This association is floristically similar to ~Amelanchier canadensis - Viburnum spp. - Morella pensylvanica Scrub Forest (CEGL006379)$$, which is a successional type.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: This association is generally a maritime forest or scrub forest, although physiognomy can vary considerably. Trees found in this community are usually stunted and flat-topped; the canopy may be only 3-7 m tall. Dominant trees vary locally but often include Prunus serotina, Sassafras albidum, and Amelanchier canadensis, with admixtures of Celtis occidentalis, Quercus velutina, Pinus rigida, Juniperus virginiana, Acer rubrum, Amelanchier stolonifera, and in southern occurrences Quercus coccinea, Quercus falcata, Liquidambar styraciflua, and/or Ilex opaca. Additional shrub species may also contribute substantially to the canopy and include Vaccinium corymbosum, Morella pensylvanica (= Myrica pensylvanica), Gaylussacia baccata, Prunus maritima, Viburnum recognitum, Viburnum dentatum, and Rosa virginiana. A true shrub layer is generally not present. Lianas are common and can be dense in the canopy or in the ground layer; species include Parthenocissus quinquefolia, Toxicodendron radicans, Smilax rotundifolia, and Smilax glauca. The understory is generally sparse with tree or vine seedlings plus herbaceous species, including Aralia nudicaulis, Moehringia lateriflora (= Arenaria lateriflora), Maianthemum stellatum (= Smilacina stellata) and Maianthemum canadense. Several invasive species can be prevalent in this association, including Lonicera morrowii, Lonicera japonica, Ligustrum vulgare, Berberis vulgaris, and Celastrus orbiculatus. The regionally uncommon species Tilia americana var. heterophylla has been reported from one occurrence (Svenson 1970).

Dynamics:  This association occupies a transitional zone between dune grasslands or low dune shrublands and maritime forest. Physiognomy is highly variable depending on degree of exposure to wind and salt spray and depending on the overall height of the surrounding dunes.

Environmental Description:  This association occurs most often on stabilized backdunes, generally leeward of secondary dunes or in protected hollows. It also occurs on bluffs or in more interior coastal areas. It is subject to varying degrees of wind and salt spray. Soils are coarse, well-drained sand subject to considerable shifting during coastal storms, or till and sand deposits of terminal moraines.

Geographic Range: The range of this community (estimated at 400 square km) is from southern New Hampshire to Delaware but is restricted to the coast.

Nations: US

States/Provinces:  CT, DE, MA, ME?, NH, NJ, NY, RI




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: G2G3

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: > High dune thicket (Martin 1959b)
>< High mesic thicket (Martin 1959b)
> High xeric thicket (Martin 1959b)
? Maritime Forest, Dune Subtype, Deciduous Forest (Rawinski 1984a)
? SNE Maritime Forest on Dunes/Maritime Juniper Forest (Rawinski 1984a)
? SNE Maritime Forest on Uplands/Maritime Oak Forest (Rawinski 1984a) [also Coastal Forest.]
? SNE coastal rocky headland community (Rawinski 1984a)
? Scrub Forest (McDonnell 1979)
? Southern New England forest on dunes (Rawinski 1984a)
>< Successional Maritime Forest (Reschke 1990)
= Successional Maritime Forest (Sneddon and Zaremba 2004)
< Sunken Forest (Art 1987)
= Sunken forest (Dunlop and Crow 1985)

Concept Author(s): L.A. Sneddon

Author of Description: S.L. Neid, L.A. Sneddon, E. Largay

Acknowledgements: No Data Available

Version Date: 03-26-07

  • Art, H. W. 1987. Patterns of community dynamics in the Sunken Forest: 1967 to 1985 and 1985 to 1986. National Park Service, North Atlantic Regional Office. 66 pp.
  • Breden, T. F. 1989. A preliminary natural community classification for New Jersey. Pages 157-191 in: E. F. Karlin, editor. New Jersey''s rare and endangered plants and animals. Institute for Environmental Studies, Ramapo College, Mahwah, NJ. 280 pp.
  • Breden, T. F., Y. R. Alger, K. S. Walz, and A. G. Windisch. 2001. Classification of vegetation communities of New Jersey: Second iteration. Association for Biodiversity Information and New Jersey Natural Heritage Program, Office of Natural Lands Management, Division of Parks and Forestry, New Jersey Department of Environmental Protection, Trenton.
  • Coxe, R. 2009. Guide to Delaware vegetation communities. Spring 2009 edition. State of Delaware, Division of Fish and Wildlife, Delaware Natural Heritage Program, Smyrna.
  • Dowhan, J. J., and R. Rozsa. 1989. Flora of Fire Island, Suffolk Country, New York. Bulletin of the Torrey Botanical Club 116:265-282.
  • Dunlop, D. A., and G. E. Crow. 1985. The vegetation and flora of the Seabrook Dunes with special reference to rare plants. Rhodora 87:471-486.
  • Eastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boston, MA.
  • Edinger, G. J., A. L. Feldmann, T. G. Howard, J. J. Schmid, E. Eastman, E. Largay, and L. A. Sneddon. 2008a. Vegetation classification and mapping at Gateway National Recreation Area. Technical Report NPS/NER/NRTR--2008/107. National Park Service, Philadelphia, PA. 283 pp.
  • Edinger, G. J., D. J. Evans, S. Gebauer, T. G. Howard, D. M. Hunt, and A. M. Olivero, editors. 2014a. Ecological communities of New York state. Second edition. A revised and expanded edition of Carol Reschke''s ecological communities of New York state. New York Natural Heritage Program, New York State Department of Environmental Conservation, Albany, NY.
  • Enser, R. W., and J. A. Lundgren. 2006. Natural communities of Rhode Island. A joint project of the Rhode Island Department of Environmental Management Natural Heritage Program and The Nature Conservancy of Rhode Island. Rhode Island Natural History Survey, Kingston. 40 pp. [www.rinhs.org]
  • Klopfer, S. D., A. Olivero, L. Sneddon, and J. Lundgren. 2002. Final report of the NPS Vegetation Mapping Project at Fire Island National Seashore. Conservation Management Institute, GIS & Remote Sensing Division, College of Natural Resources, Virginia Tech, Blacksburg, VA. 193 pp.
  • Largay, E. F., and L. A. Sneddon. 2017. Vegetation mapping and classification of Boston Harbor Islands National Recreation Area. Technical Report NPS/NER/NRTR--2017/1529. National Park Service, Fort Collins, CO.
  • Martin, W. E. 1959b. The vegetation of Island Beach State Park, New Jersey. Ecological Monographs 29:1-46.
  • McDonnell, M. J. 1979. The flora of Plum Island, Essex County, Massachusetts. University of New Hampshire, Agricultural Experiment Station. Station Bulletin No. 513. Durham, NH. 110 pp.
  • Metzler, K., and J. Barrett. 2006. The vegetation of Connecticut: A preliminary classification. State Geological and Natural History Survey, Report of Investigations No. 12. Connecticut Natural Diversity Database, Hartford, CT.
  • NRCS [Natural Resources Conservation Service]. 2001b. Soil survey of Gateway National Recreation Area, New York and New Jersey. USDA Natural Resources Conservation Service and USDI National Park Service, Gateway National Recreation Area in partnership with Cornell University Agricultural Experiment Station and New York City Soil and Water Conservation District.
  • Rawinski, T. 1984a. Natural community description abstract - southern New England calcareous seepage swamp. Unpublished report. The Nature Conservancy, Boston, MA. 6 pp.
  • Reschke, C. 1990. Ecological communities of New York State. New York Natural Heritage Program, New York State Department of Environmental Conservation, Latham, NY. 96 pp.
  • Sneddon, L. A., Zaremba, R. E., and M. Adams. 2010. Vegetation classification and mapping at Cape Cod National Seashore, Massachusetts. Natural Resources Technical Report NPS/NER/NRTR--2010/147. National Park Service, Philadelphia, PA. 481 pp. [http://biology.usgs.gov/npsveg/caco/cacorpt.pdf]
  • Sneddon, L., and R. Zaremba. 2004. Vegetation classification of Cape Cod National Seashore. U.S. Geological Survey-National Park Service Vegetation Mapping Program. 240 pp.
  • Sperduto, D. D., and W. F. Nichols. 2004. Natural communities of New Hampshire: A guide and classification. New Hampshire Natural Heritage Bureau, DRED Division of Forests and Lands, Concord. 242 pp.
  • Stalter, R. 1979. The major plant communities of the Fire Island National Seashore. Pages 177-181 in: R. M. Linn, editor. Proceedings of the first conference on Scientific Research in the National Parks. USDI National Park Service, Washington, DC.
  • Svenson, H. K. 1970. A linden (Tilia) forest on Cape Cod (with extended notes on Tilia neglecta, Bromus pubescens, and Ribes hirtellum). Rhodora 72:339-350.
  • Swain, P. C., and J. B. Kearsley. 2014. Classification of the natural communities of Massachusetts. Version 2.0. Natural Heritage & Endangered Species Program, Massachusetts Division of Fisheries and Wildlife. Westborough, MA. [http://www.mass.gov/nhesp/http://www.mass.gov/eea/agencies/dfg/dfw/natural-heritage/natural-communities/classification-of-natural-communities.html]