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CEGL004418 Liquidambar styraciflua - Liriodendron tulipifera / Lindera benzoin / Arisaema triphyllum Floodplain Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Sweetgum - Tuliptree / Northern Spicebush / Jack-in-the-Pulpit Floodplain Forest

Colloquial Name: Upper Southeast Sweetgum - Tuliptree Small Stream Forest

Hierarchy Level:  Association

Type Concept: These low-elevation forests develop along relatively acidic soils on small streams in the Coastal Plain of Maryland and Virginia, extending west across the Virginia and North Carolina Piedmont to the Cumberland Plateau and Ridge and Valley. The topographic features of floodplains can heavily influence the makeup of individual examples of this association. The canopy, subcanopy, shrub, and herbaceous layers often are well-developed. Dominant canopy species always include Liquidambar styraciflua and Liriodendron tulipifera, while Acer floridanum (in the eastern part of the range), Platanus occidentalis, and Acer rubrum var. rubrum may also make up significant amounts of the canopy. This community type exists as a continuum between two subtypes, i.e., the tuliptree subtype and the sweetgum subtype. In some examples, only one or the other dominates the canopy, but in many examples, both are equally dominant. Common species in the canopy and understory include Ilex opaca var. opaca, Aesculus sylvatica, Betula nigra, Carpinus caroliniana ssp. caroliniana, Cornus florida, Carya cordiformis, Fagus grandifolia, Fraxinus americana, Fraxinus pennsylvanica, Halesia tetraptera var. tetraptera, Juglans nigra, Juniperus virginiana var. virginiana, Morus rubra var. rubra, Nyssa sylvatica, Ostrya virginiana, Oxydendrum arboreum, Pinus echinata, Prunus serotina var. serotina, Quercus alba, Quercus rubra var. rubra, Ulmus rubra, Ulmus americana, and Ulmus alata. Euonymus americanus, Asimina triloba, Lindera benzoin var. benzoin, and Corylus americana are common in the shrub layer. The herbaceous layer is species-rich and often has good sedge development. The exotics Microstegium vimineum, Glechoma hederacea, Rosa multiflora, Ligustrum sinense, and Lonicera japonica are common in this community.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Low-quality occurrences of this type may look very similar to some occurrences of ~Liquidambar styraciflua - (Liriodendron tulipifera) Ruderal Wet Forest (CEGL007330)$$. The presence of higher quality patches of native herbs and stands of native shrubs such as Lindera benzoin is the best way to distinguish these two types. In addition, stands of CEGL007330 will generally be more even-aged and single species-dominated than this association (CEGL004418).

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: The canopy, subcanopy, shrub, and herbaceous layers of stands of this association are often well-developed. Dominant canopy species always include Liquidambar styraciflua and Liriodendron tulipifera, while Acer floridanum (= Acer barbatum) (in the southern part of the range), Platanus occidentalis, and Acer rubrum var. rubrum may also make up significant amounts of the canopy. This community type exists as a continuum between two subtypes, i.e., the tuliptree subtype and the sweetgum subtype. In some examples, only one or the other dominates the canopy. However, in many examples, both are equally dominant. Other common species in the canopy and understory include Ilex opaca var. opaca, Aesculus sylvatica, Carpinus caroliniana ssp. caroliniana, Carya cordiformis, Cornus florida, Fagus grandifolia, Juglans nigra, Betula nigra, Morus rubra var. rubra, Ostrya virginiana (= var. virginiana), Oxydendrum arboreum, Pinus echinata, Prunus serotina var. serotina, Quercus alba, Quercus rubra var. rubra, Ulmus rubra, Ulmus americana, Ulmus alata, Juniperus virginiana var. virginiana, Nyssa sylvatica, Fraxinus americana, Halesia tetraptera var. tetraptera, Arundinaria gigantea (= ssp. gigantea), and Fraxinus pennsylvanica. Euonymus americanus, Lindera benzoin var. benzoin, and Corylus americana are common and dominant in the shrub layer. Other shrub species that may be present include Viburnum acerifolium, Viburnum nudum var. nudum, Viburnum prunifolium, Viburnum rufidulum, Hamamelis virginiana, Asimina triloba, and Ilex decidua, among others. On the most acidic sites of the Maryland Coastal Plain, Clethra alnifolia, Vaccinium corymbosum, and Magnolia virginiana may be present. Vines are prominent and include Vitis rotundifolia, Apios americana, Campsis radicans, Aristolochia macrophylla, Bignonia capreolata, Dioscorea quaternata, Gelsemium sempervirens, Parthenocissus quinquefolia (= var. quinquefolia), Passiflora lutea, Smilax bona-nox, Smilax glauca, Smilax hugeri, Smilax rotundifolia, and Toxicodendron radicans ssp. radicans. The herbaceous layer is species-rich and often has good sedge development. Common species in this layer include Thalictrum thalictroides, Trillium cuneatum, Arisaema triphyllum, Asplenium platyneuron var. platyneuron, Botrychium virginianum, Carex spp., Carex impressinervia, Carex striatula, Cinna arundinacea, Collinsonia canadensis, Deparia acrostichoides, Dichanthelium clandestinum, Elymus virginicus, Eurybia divaricata, Galium circaezans, Geum canadense, Medeola virginiana, Packera aurea, Poa alsodes, Polygonatum pubescens, Polystichum acrostichoides, Rudbeckia laciniata, Scutellaria integrifolia, Symphyotrichum prenanthoides, and Viola striata. Thelypteris noveboracensis is a common patch-dominant in the northern part of the range and the Uwharrie Mountains of North Carolina. Carex kraliana, which is evidently near the northern limit of its (primarily southeastern United States) range and is presently known from only two Maryland sites, has been found in this vegetation type at Thomas Stone NHS. A specimen collected at Thomas Stone NHS was cited as a paratype in the description of the species (Naczi et al. 2002). The exotics Microstegium vimineum, Glechoma hederacea, Rosa multiflora, Ligustrum sinense, and Lonicera japonica are common in this community. Other exotics that colonize quickly in disturbed and fragmented versions of this association include Wisteria sinensis, Clematis terniflora, Hedera helix, and Elaeagnus sp.

Dynamics:  The topographic features characteristic of larger-stream floodplains are poorly differentiated in the small-stream habitats of this type, and vegetation zonation is absent to poorly defined. It is unclear what processes create pure sweetgum versus pure tuliptree forests, but dominance is most likely a factor of seed source and amount of flooding (sweetgum may be able to tolerate higher water levels than tuliptree). ~Liquidambar styraciflua - (Liriodendron tulipifera) Ruderal Wet Forest (CEGL007330)$$ is very similar to this community but is usually less than 40 years old and very even-aged with a heavy infestation of invasive exotics and fewer native species in the shrub and herb layer.

Environmental Description:  These forests develop along small streams or in small patches on river floodplains. Soils are relatively acidic and relatively well-drained. Topographic differences from one floodplain to another, such as gradient and height above the creek, as well as floodplain microtopography (i.e., depositional landforms such as natural levees and sloughs) may influence the variation of vegetation within this association. However, in most floodplains supporting this type, the distinct alluvial landforms are poorly developed or occur at very small scales. At some sites, evidence of infrequent flooding includes fluvial topography, mixed exposed sand, light flotsam and leaf litter accumulations, and lack of soil horizon development.

Geographic Range: This association is found in the Chesapeake Bay Lowlands, the Piedmont, and other low-elevation interior ecoregions (e.g., parts of the Cumberland Plateau and Ridge and Valley) from Maryland and West Virginia south to the Carolinas and west into Tennessee. It is defined as being absent from the Mid-Atlantic Coastal Plain of southeastern Virginia, the Carolinas, and Georgia. Its status in the Upper East Gulf Coastal Plain is unknown.

Nations: US

States/Provinces:  DC, GA, MD, NC, SC?, TN, VA, WV




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: G4

Greasons: No Data Available


Concept Lineage: Liriodendron tulipifera - Acer rubrum - Liquidambar styraciflua / Medeola virginiana Forest (CEGL006601) merged into this type (CEGL004418). The the two types were overlapping in concept and were merged. CEGL007327 merged into CEGL006601 which was subsequently merged into CEGL004418.

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Liquidambar styraciflua - Liriodendron tulipifera / Lindera benzoin Temporarily Flooded Forest (McCoy and Fleming 2000)
? Liquidambar styraciflua - Quercus palustris / Carpinus caroliniana / Carex intumescens Forest (Meininger and McCarthy 1998)
= Liquidambar styraciflua / Lindera benzoin / Arisaema triphyllum ssp. triphyllum Temporarily Flooded Forest (Coulling 1999)
= Liriodendron tulipifera - Fagus grandifolia / Thelypteris noveboracensis Forest (Fleming and Patterson 2003)
< Maple-Gum Association of the Western Shore District (Shreve et al. 1910)

Concept Author(s): R.K. Peet

Author of Description: R.K. Peet, R. White, M. Pyne, G.P. Fleming, S.C. Gawler

Acknowledgements: No Data Available

Version Date: 01-10-18

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