Print Report

CEGL000471 Thuja plicata / Aralia nudicaulis Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Western Red-cedar / Wild Sarsaparilla Forest

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This is a late-seral evergreen conifer forest found in the mid elevations in the Okanogan Highlands east of the Kettle Mountain crest in Washington and British Columbia, with outliers in Montana. It appears on stream terraces representing xero-riparian conditions and on lower slopes and benches, mostly below 1065 m (3500 feet) in Washington and 1220 m (4000 feet) in Montana. Its name reflects a potential vegetation. The existing vegetation is dominated by a closed canopy of Picea engelmannii, Thuja plicata, and Pseudotsuga menziesii trees. The undergrowth can contain scattered tall deciduous shrubs such as Acer glabrum and Alnus viridis ssp. sinuata. The short and dwarf-shrub layers have both greater diversity and cover than the tall-shrub layer, with Linnaea borealis and Cornus canadensis being the most constant and having appreciable coverages. The herbaceous component is characterized by a rich assortment of mesic site forbs, such as Aralia nudicaulis, Clintonia uniflora, Galium triflorum, Maianthemum stellatum, Tiarella trifoliata, and Prosartes hookeri; graminoids, with the exception of the highly constant Bromus vulgaris (or Bromus ciliatus), are a minor element.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Currently there are at least two conceptions of this type, one is that of a potential vegetation type identified by the mere presence and successful reproduction of Thuja plicata (Pfister et al. 1977, Williams et al. 1995); Williams et al. (1995) also recognize the type based on at least 10% cover of Thuja and less than 10% cover of Tsuga heterophylla. A modified cover type approach (as encouraged by the USNVC) is reflected by the approach taken in Waterton-Glacier International Peace Park (IPP); Thuja must comprise at least 25% of the cover of the upper canopy and Tsuga less than 25% of this stratum. Under the potential-vegetation concept a considerable number of seral tree species are capable of dominating the site, whereas with the IPP concept, primarily mature to late-seral and climax stages are recognized. In Montana this association was formerly recognized as a phase of the Thuja plicata / Clintonia uniflora Habitat Type (Pfister et al. 1977), but with accumulating compelling information linking this phase with particular environmental parameters, it is appropriate to recognize it at the association level, as exemplified by Williams et al. (1995). However, following intensive field sampling in IPP, it is deemed prudent to recognize Aralia nudicaulis as an indicator only at 1% or greater canopy cover; this contrasts with Pfister et al. (1977) and Cooper et al. (1987) who recognize its mere presence not confined to microsites as indicative. In contrast Williams et al. (1995) stipulate that 5% be present to recognize the type. Pfister et al. (1977) and Williams et al. (1995) also use Gymnocarpium dryopteris, Athyrium filix-femina, Asarum caudatum, Actaea rubra, and Cornus canadensis as alternative indicators for this association. All of these species are also used as indicators of other plant associations (both with and without Thuja as overstory indicator), and the first three are clearly indicative of more mesic environments or at least a more maritime climatic regime than reflected by the presence of Aralia nudicaulis.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: The existing vegetation is dominated by a relatively closed canopy of Thuja plicata, Picea engelmannii, and Pseudotsuga menziesii; throughout all but the easternmost extent of the type, Abies grandis often contributes the greatest canopy cover. The undergrowth can contain scattered tall deciduous shrubs, such as Acer glabrum and Amelanchier alnifolia. The short-shrub layer is often moderately dense, though not highly diverse with the most constant species being Rubus parviflorus, Symphoricarpos albus, Spiraea betulifolia, and Rosa gymnocarpa. In the dwarf-shrub layer Chimaphila umbellata, Linnaea borealis, and Cornus canadensis have the highest constancy and cover. A rich assortment of mesic-site forbs consistently includes Aralia nudicaulis, Asarum caudatum, Clintonia uniflora, Prosartes hookeri (= Disporum hookeri), Galium triflorum, Gymnocarpium dryopteris, Maianthemum stellatum, Tiarella trifoliata, Trillium ovatum, and Viola orbiculata. Bromus vulgaris is the only graminoid with greater than 50% constancy. Though some would include stands with relatively high cover of Gymnocarpium dryopteris, Athyrium filix-femina, Asarum caudatum, and Cornus canadensis in this type, from all indications these species would reflect other, more mesic environments (and types).

Dynamics:  Some stands of ~Pinus contorta / Shepherdia canadensis Forest (CEGL000163)$$ occur on sites environmentally similar to those supporting ~Thuja plicata / Aralia nudicaulis Forest (CEGL000471)$$ leading Williams et al. (1995) to speculate that such areas have burned intensively one or more times. Subsequently, nutrient pools and organic matter concentrations have declined such that sites have not fully recovered. Similar vegetation patterns are expressed on sites that have been cleared as fields due to gentle slopes, low elevations (relatively warm) and deep soils; Pinus contorta usually colonizes these abandoned homesteads and fields (contrary to the course of succession following natural disturbance, i.e., patchy underburns or spotty crown fires).

Environmental Description:  This is a late-seral evergreen conifer forest found in the mid elevations in the Okanogan Highlands east of the Kettle Mountain crest in Washington and British Columbia, with outliers in northwestern Montana as far east as the Continental Divide. It appears on stream terraces representing xero-riparian conditions and on lower slopes and benches, mostly below 1070 m (3500 feet) in Washington and 1220 m (4000 feet) in Montana. Sites are postulated to be relatively warm and moist. Soils have fine-textured upper horizons, often with a high concentration of volcanic ash, overlying coarse-textured alluvium or glacial outwash and drift; thus soils are expected to be nutrient-rich and well-drained.

Geographic Range: This association is restricted to the eastern portion of the Okanogan Highlands and penetrates into northwestern Montana extending as far east as the Continental Divide (beyond which Thuja plicata does not extend); it has not been recognized for northern Idaho, but this may be due to the omnipresence of Tsuga heterophylla, which was given precedence, even in trace amounts in the seedling layer, over Thuja in potential-based vegetation keys.

Nations: CA?,US

States/Provinces:  BC?, ID?, MT, WA




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: G2

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: >< Thuja plicata / Aralia nudicaulis Plant Association (Williams et al. 1995)
>< Thuja plicata / Clintonia uniflora Habitat Type, Aralia nudicaulis Phase (Pfister et al. 1977)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Aralia nudicaulis Phase (Pfister et al. 1977)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Aralia nudicaulis Phase (Cooper et al. 1987)

Concept Author(s): R.C. Crawford and S.V. Cooper

Author of Description: R.C. Crawford and S.V. Cooper

Acknowledgements: No Data Available

Version Date: 04-14-04

  • Bourgeron, P. S., and L. D. Engelking, editors. 1994. A preliminary vegetation classification of the western United States. Unpublished report. The Nature Conservancy, Western Heritage Task Force, Boulder, CO. 175 pp. plus appendix.
  • Cooper, S. V., K. E. Neiman, R. Steele, and D. W. Roberts. 1987. Forest habitat types of northern Idaho: A second approximation. General Technical Report INT-236.USDA Forest Service, Intermountain Research Station, Ogden, UT. 135 pp. [reprinted in 1991]
  • Hansen, P. L., R. D. Pfister, K. Boggs, B. J. Cook, J. Joy, and D. K. Hinckley. 1995. Classification and management of Montana''s riparian and wetland sites. Miscellaneous Publication No. 54. Montana Forest and Conservation Experiment Station, School of Forestry, University of Montana. 646 pp. plus posters.
  • Hop, K., M. Reid, J. Dieck, S. Lubinski, and S. Cooper. 2007. U.S. Geological Survey-National Park Service Vegetation Mapping Program: Waterton-Glacier International Peace Park. U.S. Geological Survey, Upper Midwest Environmental Sciences Center, La Crosse, WI. 131 pp. plus Appendices A-L.
  • Kovalchik, B. L. 1993. Riparian plant associations on the national forests of eastern Washington - Draft version 1. USDA Forest Service, Colville National Forest, Colville, WA. 203 pp.
  • MTNHP [Montana Natural Heritage Program]. 2002b. List of ecological communities for Montana. Montana Natural Heritage Program, Montana State Library, Helena, MT.
  • Pfister, R. D., B. L. Kovalchik, S. F. Arno, and R. C. Presby. 1977. Forest habitat types of Montana. General Technical Report INT-34. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 174 pp.
  • Reid, M. S., S. V. Cooper, and G. Kittel. 2004. Vegetation classification of Waterton-Glacier International Peace Park. Final report for USGS-NPS Vegetation Mapping Program, International Peace Park Mapping Project. NatureServe, Arlington VA.
  • WNHP [Washington Natural Heritage Program]. 2018. Unpublished data files. Washington Natural Heritage Program, Department of Natural Resources, Olympia, WA.
  • Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.
  • Williams, C. K., B. F. Kelly, B. G. Smith, and T. R. Lillybridge. 1995. Forest plant associations of the Colville National Forest. General Technical Report PNW-GTR-360. USDA Forest Service, Pacific Northwest Region, Portland, OR. 140 pp.
  • Williams, C. K., T. R. Lillybridge, and B. G. Smith. 1990b. Forested plant associations of the Colville National Forest. Report prepared for USDA Forest Service, Colville National Forest, Colville, WA. 133 pp.
  • Williams, C. K., and T. R. Lillybridge. 1985. Forested plant associations of the Colville National Forest. Draft. Unpublished field guide prepared for USDA Forest Service.