Print Report
CEGL006057 Quercus montana - Quercus rubra / Hamamelis virginiana Forest
Type Concept Sentence: This dry-mesic chestnut oak - northern red oak forest of the Central Appalachians is intermediate both in composition and habitat between dry ridgetop oak forests and richer oak - hickory forests in coves or on more sheltered slopes; the tree canopy is codominated by Quercus montana and Quercus rubra, with Liriodendron tulipifera, Acer rubrum, Carya spp., Acer saccharum, Tilia americana, Fagus grandifolia, and Betula lenta.
Common (Translated Scientific) Name: Chestnut Oak - Northern Red Oak / American Witch-hazel Forest
Colloquial Name: Central Appalachian Dry-Mesic Chestnut Oak - Northern Red Oak Forest
Hierarchy Level: Association
Type Concept: This closed-canopy, dry-mesic oak forest of the central Appalachian Mountains is a montane forest of somewhat protected rocky slopes. The canopy is codominated by Quercus montana and Quercus rubra. Associated canopy species include Liriodendron tulipifera, Acer rubrum, Carya glabra, Carya ovalis, Carya tomentosa, Acer saccharum, Tilia americana, Fagus grandifolia, and Betula lenta. The tall-shrub layer is most often characterized by Hamamelis virginiana and Acer pensylvanicum. The lower shrub layer is patchy and contains a mixture of scrambling vines, ericads, and non-ericaceous species. The herbaceous layer is usually sparse but may include Dryopteris marginalis, Dioscorea quaternata, Eurybia divaricata, Ageratina altissima, Polygonatum biflorum, Solidago caesia, Festuca subverticillata, Thelypteris noveboracensis, Sanicula trifoliata, Prenanthes altissima, Polystichum acrostichoides, Desmodium nudiflorum, Galium latifolium, Houstonia purpurea, and Maianthemum racemosum. This association is more or less intermediate in site conditions and composition between oak / heath forests of exposed, xeric, infertile sites and richer cove or montane oak-hickory forests of sheltered, fertile sites.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: No Data Available
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: The vegetation is usually a closed-canopy forest codominated by Quercus montana (= Quercus prinus) and Quercus rubra in variable proportions. Over the full geographic range, overstory associates are reported to include Liriodendron tulipifera, Fraxinus americana, Tilia americana, Betula lenta, Acer rubrum, Magnolia acuminata, Nyssa sylvatica, Robinia pseudoacacia, Carya glabra, Carya ovalis, and Carya tomentosa (= Carya alba). Less frequent, and more local, overstory and understory trees include Acer saccharum, Amelanchier arborea, Asimina triloba, Fagus grandifolia, Ostrya virginiana, and Tsuga canadensis. A tall-shrub layer is occasionally absent but usually characterized by Hamamelis virginiana and, less frequently, by Cornus florida and Acer pensylvanicum, the latter more common at higher elevations. The lower shrub layer contains scrambling or climbing vines of Parthenocissus quinquefolia, Vitis aestivalis, and Toxicodendron radicans, along with Viburnum acerifolium, Hydrangea arborescens, Vaccinium pallidum, and Vaccinium stamineum. In general, ericaceous species are patchy to sparse in this community. The herbaceous layer is usually sparse but may include Dryopteris marginalis, Dioscorea quaternata, Eurybia divaricata (= Aster divaricatus), Ageratina altissima, Polygonatum biflorum, Solidago caesia, Festuca subverticillata, Thelypteris noveboracensis, Sanicula trifoliata, Prenanthes altissima, Polystichum acrostichoides, Desmodium nudiflorum, Galium latifolium, Houstonia purpurea, and Maianthemum racemosum. Although not one of the more constant herbs, Aralia nudicaulis may occasionally dominate the herb layer of this community in large, clonal patches. This association is more-or-less intermediate in site conditions and composition between oak/heath forests of exposed, xeric, infertile sites and richer cove or montane oak-hickory forests of sheltered, fertile sites.
Dynamics: Wind and ice damage to tree crowns, damage to Cornus florida from dogwood anthracnose (Discula destructiva), and a few small fire scars were disturbances noted in plots. Although Castanea dentata logs and wood were not abundant in plots, frequent sprouts indicate that Castanea dentata was at least an associate tree in this type prior to the arrival of chestnut blight. The northwest slopes of Peters Mountain in Alleghany County, Virginia, contains old-growth examples of the type with large, widely spaced canopy trees in the 43- to 72-cm (17-28 inches) dbh range. Representative old-age trees include a 59-cm (23 inches) dbh Quercus montana >220 years old; a 63-cm (25 inches) dbh Quercus montana about179 years old; a 67-cm (26 inches) dbh Quercus montana 265 years old; and a 71-cm (28 inches) dbh Quercus rubra >247 years old (Fleming and Moorhead 2000).
Environmental Description: Sites occupied by this dry-mesic oak forest are mostly protected rocky mountain slopes. In the Central Appalachians of Virginia, West Virginia, and Maryland, the type occurs at low and middle elevations, from <300 m (1000 feet) to about 1100 m (3600 feet), reaching optimal development at 610-915 m (2000-3000 feet). Habitats are underlain by a variety of bedrock types, including metabasalt (greenstone), pyroxene-rich granitic rocks, Antietam and Tuscarora quartzites, metasiltstone and phyllite, shale, and sedimentary material (interbedded sandstone, siltstone, and shale). Among plot-sampled Mid-Atlantic stands, lower to middle slope topographic positions predominate, along with steep (mean = 27°), usually concave slopes, and relatively high surface cover of outcrops, boulders, and stones. Slope aspect is variable, but the majority of aspects range from north to southeast. Soil samples collected from plots were strongly to very strongly acidic (mean pH = 4.8) but had moderately high levels of calcium (mean = 1019 ppm), reflecting the frequent occurrence of this community on moderately base-rich substrates.
Geographic Range: This association occurs throughout the Central Appalachian region of Virginia, West Virginia, Maryland, Pennsylvania, and possibly farther north. In Virginia, it is a large-patch community type in both the Northern Blue Ridge and Ridge and Valley provinces. Small-patch outliers of this type occur in rocky, sheltered ravines of the northern Virginia and Maryland Piedmont.
Nations: US
States/Provinces: DE, MD, NJ, PA, VA, WV
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.683617
Confidence Level: Moderate
Confidence Level Comments: No Data Available
Grank: G5
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.2 Cool Temperate Forest & Woodland Formation | F008 | 1.B.2 |
Division | 1.B.2.Na Eastern North American Forest & Woodland Division | D008 | 1.B.2.Na |
Macrogroup | 1.B.2.Na.2 Appalachian-Northeastern Oak - Hardwood - Pine Forest & Woodland Macrogroup | M502 | 1.B.2.Na.2 |
Group | 1.B.2.Na.2.c White Oak - Chestnut Oak - Pignut Hickory Forest & Woodland Group | G650 | 1.B.2.Na.2.c |
Alliance | A4435 Chestnut Oak / Mountain Laurel - Blue Ridge Blueberry Forest Alliance | A4435 | 1.B.2.Na.2.c |
Association | CEGL006057 Chestnut Oak - Northern Red Oak / American Witch-hazel Forest | CEGL006057 | 1.B.2.Na.2.c |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: > Quercus montana - Quercus rubra / Acer pensylvanicum - Hamamelis virginiana Forest (Fleming and Moorhead 2000)
= Quercus montana - Quercus rubra / Cornus florida / Viburnum acerifolium Forest (Young et al. 2007a)
= Quercus montana - Quercus rubra / Vitis aestivalis var. bicolor - Parthenocissus quinquefolia / Aralia nudicaulis - Dryopteris marginalis Forest (Fleming 2002b)
> Quercus montana - Robinia pseudoacacia / Ribes rotundifolium Association (Rawinski et al. 1994)
= Quercus prinus - Quercus rubra / Hamamelis virginiana Forest (Fleming and Coulling 2001)
= Quercus prinus - Quercus rubra / Hamamelis virginiana Forest (Fleming 2007)
> Quercus rubra - Magnolia acuminata Association (Fleming and Moorhead 1996)
= Quercus rubra - Quercus montana / Parthenocissus quinquefolia - Vitis aestivalis var. bicolor / Dryopteris marginalis - Aralia nudicaulis Forest (Fleming and Patterson 2003)
= Quercus rubra - Quercus montana / Parthenocissus quinquefolia / Aralia nudicaulis - Dryopteris marginalis Forest (Fleming 2002a)
> Quercus rubra - Quercus prinus - Liriodendron tulipifera / Parthenocissus quinquefolia - Dryopteris marginalis Association (Rawinski et al. 1996)
= Chestnut Oak - Black Birch community (Ehrenfeld 1977)
< Chestnut Oak: 44 (Eyre 1980) [chestnut oak - northern red oak variant]
> Chestnut oak-red oak/ericad forest: (matrix) N slopes (CAP pers. comm. 1998)
? Mixed oak / hardwoods mesic forest (Windisch 1993)
? Red Oak - Chestnut Oak Community Type (Stephenson and Adams 1991)
= Quercus montana - Quercus rubra / Cornus florida / Viburnum acerifolium Forest (Young et al. 2007a)
= Quercus montana - Quercus rubra / Vitis aestivalis var. bicolor - Parthenocissus quinquefolia / Aralia nudicaulis - Dryopteris marginalis Forest (Fleming 2002b)
> Quercus montana - Robinia pseudoacacia / Ribes rotundifolium Association (Rawinski et al. 1994)
= Quercus prinus - Quercus rubra / Hamamelis virginiana Forest (Fleming and Coulling 2001)
= Quercus prinus - Quercus rubra / Hamamelis virginiana Forest (Fleming 2007)
> Quercus rubra - Magnolia acuminata Association (Fleming and Moorhead 1996)
= Quercus rubra - Quercus montana / Parthenocissus quinquefolia - Vitis aestivalis var. bicolor / Dryopteris marginalis - Aralia nudicaulis Forest (Fleming and Patterson 2003)
= Quercus rubra - Quercus montana / Parthenocissus quinquefolia / Aralia nudicaulis - Dryopteris marginalis Forest (Fleming 2002a)
> Quercus rubra - Quercus prinus - Liriodendron tulipifera / Parthenocissus quinquefolia - Dryopteris marginalis Association (Rawinski et al. 1996)
= Chestnut Oak - Black Birch community (Ehrenfeld 1977)
< Chestnut Oak: 44 (Eyre 1980) [chestnut oak - northern red oak variant]
> Chestnut oak-red oak/ericad forest: (matrix) N slopes (CAP pers. comm. 1998)
? Mixed oak / hardwoods mesic forest (Windisch 1993)
? Red Oak - Chestnut Oak Community Type (Stephenson and Adams 1991)
- Breden, T. F., Y. R. Alger, K. S. Walz, and A. G. Windisch. 2001. Classification of vegetation communities of New Jersey: Second iteration. Association for Biodiversity Information and New Jersey Natural Heritage Program, Office of Natural Lands Management, Division of Parks and Forestry, New Jersey Department of Environmental Protection, Trenton.
- CAP [Central Appalachian Forest Working Group]. 1998. Central Appalachian Working group discussions. The Nature Conservancy, Boston, MA.
- Coxe, R. 2009. Guide to Delaware vegetation communities. Spring 2009 edition. State of Delaware, Division of Fish and Wildlife, Delaware Natural Heritage Program, Smyrna.
- Eastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boston, MA.
- Ehrenfeld, J. G. 1977. Vegetation of Morristown National Historical Park: Ecological analysis and management alternatives. Final Report. USDI National Park Service Contract No. 1600-7-0004. 166 pp.
- Eyre, F. H., editor. 1980. Forest cover types of the United States and Canada. Society of American Foresters, Washington, DC. 148 pp.
- Fike, J. 1999. Terrestrial and palustrine plant communities of Pennsylvania. Pennsylvania Natural Diversity Inventory. Pennsylvania Department of Conservation and Recreation, Bureau of Forestry, Harrisburg, PA. 86 pp.
- Fleming, G. P. 2002a. Ecological communities of the Bull Run Mountains, Virginia: Baseline vegetation and floristic data for conservation planning and natural area stewardship. Natural Heritage Technical Report 02-12. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 274 pp. plus appendices.
- Fleming, G. P. 2002b. Preliminary classification of Piedmont & Inner Coastal Plain vegetation types in Virginia. Natural Heritage Technical Report 02-14. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 29 pp.
- Fleming, G. P. 2007. Ecological communities of the Potomac Gorge in Virginia: Composition, floristics, and environmental dynamics. Natural Heritage Technical Report 07-12. Unpublished report submitted to the National Park Service. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 341 pp. plus appendices.
- Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
- Fleming, G. P., and K. D. Patterson. 2003. Preliminary vegetation classification for the National Capitol Region parks. Regional (VA-WVA-MD-DC) analysis prepared for NatureServe and USGS-NPS Vegetation Mapping Program, March 2003. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
- Fleming, G. P., and K. D. Patterson. 2009a. A vegetation classification for the Appalachian Trail: Virginia south to Georgia. Virginia Department of Conservation and Recreation, Division of Natural Heritage. In-house analysis, March 2009.
- Fleming, G. P., and K. D. Patterson. 2009b. Classification of selected Virginia montane wetland groups. In-house analysis, December 2009. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
- Fleming, G. P., and K. Taverna. 2006. Vegetation classification for the National Capitol Region parks, western region. Regional (VA-WVA-MD-DC) analysis prepared for NatureServe and USGS-NPS Vegetation Mapping Program, March 2006. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
- Fleming, G. P., and P. P. Coulling. 2001. Ecological communities of the George Washington and Jefferson national forests, Virginia. Preliminary classification and description of vegetation types. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. 317 pp.
- Fleming, G. P., and W. H. Moorhead, III. 1996. Ecological land units of the Laurel Fork Area, Highland County, Virginia. Natural Heritage Technical Report 96-08. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 114 pp. plus appendices.
- Fleming, G. P., and W. H. Moorhead, III. 2000. Plant communities and ecological land units of the Peter''s Mountain area, James River Ranger District, George Washington and Jefferson national forests, Virginia. Natural Heritage Technical Report 00-07. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. Unpublished report submitted to the USDA Forest Service. 195 pp. plus appendices.
- Harrison, J. W. 2011. The natural communities of Maryland: 2011 working list of ecological community groups and community types. Unpublished report. Maryland Department of Natural Resources, Wildlife and Heritage Service, Natural Heritage Program, Annapolis. 33 pp.
- Harrison, J. W., compiler. 2004. Classification of vegetation communities of Maryland: First iteration. A subset of the International Classification of Ecological Communities: Terrestrial Vegetation of the United States, NatureServe. Maryland Natural Heritage Program, Maryland Department of Natural Resources, Annapolis. 243 pp.
- Lea, C. 2003. Vegetation types in the National Capital Region Parks. Draft for review by NatureServe, Virginia Natural Heritage, West Virginia Natural Heritage, Maryland Natural Heritage, and National Park Service. March 2003. 140 pp.
- Lea, C. 2004. Draft vegetation types in National Capital Region Parks. Edited by S.C. Gawler and J. Teague. Working draft for review by NatureServe, Virginia Natural Heritage, West Virginia Natural Heritage, Maryland Natural Heritage, and National Park Service. July 2004. 157 pp.
- Rawinski, T. J., G. P. Fleming, and F. V. Judge. 1994. Forest vegetation of the Ramsey''s Draft and Little Laurel Run Research Natural Areas, Virginia: Baseline ecological monitoring and classification. Natural Heritage Technical Report 94-14. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 45 pp. plus appendices.
- Rawinski, T. J., K. N. Hickman, J. Waller-Eling, G. P. Fleming, C. S. Austin, S. D. Helmick, C. Huber, G. Kappesser, F. C. Huber, Jr., T. Bailey, and T. K. Collins. 1996. Plant communities and ecological land units of the Glenwood Ranger District, George Washington and Jefferson national forests, Virginia. Virginia Department of Conservation and Recreation, Division of Natural Heritage. Natural Heritage Technical Report 96-20. Richmond. 65 pp. plus appendices.
- Sneddon, L., R. E. Zaremba, E. Largay, G. Podniesinski, S. Perles, and J. Thompson. 2008. Vegetation classification and mapping of Morristown National Historical Park, New Jersey. Technical Report NPS/NER/NRTR--2008/116. National Park Service, Philadelphia, PA. 162 pp. [http://biology.usgs.gov/npsveg/morr/morrrpt.pdf]
- Stephenson, S. L., and H. S. Adams. 1991. Upland oak forests of the Ridge and Valley Province in southwestern Virginia. Virginia Journal of Science 42:371-380.
- Vanderhorst, J. 2000b. Plant communities of Harper''s Ferry National Historical Park: Analysis, characterization, and mapping. West Virginia Natural Heritage Program, West Virginia Division of Natural Resources, Elkins, WV. 37 pp.
- WVNHP [West Virginia Natural Heritage Program]. No date. Unpublished data. West Virginia Natural Heritage Program, Elkins.
- Windisch, A. G. 1993. Natural community inventory of Picatinny Arsenal, New Jersey. Unpublished report prepared for Picatinny Arsenal, U.S. Department of Defense. The Nature Conservancy, Eastern Heritage Task Force, Trenton, NJ.
- Young, J., G. Fleming, P. Townsend, and J. Foster. 2006. Vegetation of Shenandoah National Park in relation to environmental gradients. Final Report (v.1.1). Research technical report prepared for USDI, National Park Service. USGS/NPS Vegetation Mapping Program. 92 pp. plus appendices.
- Young, J., G. Fleming, P. Townsend, and J. Foster. 2007a. Vegetation of Shenandoah National Park in relation to environmental gradients. Final Report, volume 1.1. Unpublished report submitted to the U.S. Department of the Interior, National Park Service. 103 pp. plus appendices and GIS products.
- Young, J., G. Fleming, W. Cass, and C. Lea. 2009. Vegetation of Shenandoah National Park in relation to environmental gradients, Version 2.0. Technical Report NPS/NER/NRTR--2009/142. National Park Service, Philadelphia, PA. 389 pp.