Cooperative Research Units
Education, Research And Technical Assistance For Managing Our Natural Resources
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Dr. Patrick Jodice

Research Publications

  • Lawson, A.J., Clinton T. Moore, Thomas R. Rainwater, Frances M. Nilsen, Philip M. Wilkinson, Russell H. Lowers, Louis J. Guillette Jr., K.W. McFadden, and Patrick G.R. Jodice. 2020. Nonlinear patterns in mercury bioaccumulation in American alligators are a function of predicted age. Science of the Total Environment 707:135103. <a href=""></a>
  • Zaluski, S., L.M. Soanes, J.A. Bright, A. George, P.G.R. Jodice, K. Meyer, N. Woodfield-Pascoe, J.A. Green. 2019. Potential threats facing a globally important population of the magnificent frigatebird. Tropical Zoology 32: 188-201. /doi/full/10.1080/03946975.2019.1682352
  • Lamb, J.S.*, Y.G. Satge^, P.G.R. Jodice. Environmental and behavioral drivers of annual-cycle habitat selection in a nearshore seabird, the Brown Pelican. Diversity and Distributions.
  • Satgé, Y.G., E. Rupp, and P.G.R. Jodice. 2019. A preliminary report of ongoing research of the ecology of Black-capped Petrel (Pterodroma hasitata) in Sierra de Bahoruco, Dominican Republic – I: GPS tracking of breeding adults. Unpublished Report, South Carolina Cooperative Research Unit, Clemson University, Clemson, South Carolina, USA.
  • Ferguson, L.M., Y. Satge, J. Tavano, and P.G.R. Jodice. 2018. Seabird colony registry and atlas for the Southeastern U.S. Final Report for U.S. Fish and Wildlife Service. South Carolina Cooperative Fish and Wildlife Research Unit, Clemson, SC.
  • Lamb, J.S., C.V. Fiorello, Y.G. Satge, K. Mills-Parker, M. Ziccardi, P.G.R. Jodice. 2018. Movement patterns of California brown pelicans following oiling and rehabilitation. Marine Pollution Bulletin 131:22-31
  • Loeb, S.C., P.G.R. Jodice. Activity of southeastern bats along sandstone cliffs used for rock climbing. Journal of Fish and Wildlife Management 9:255-265; e1944-687x.
  • Lamb, J.S.*, D.J. Newstead, L.M. Koczur, B.M. Ballard, C.M. Green, P.G.R. Jodice. 2018. A bridge between oceans: Overland migration of marine birds in a wind energy corridor. Journal Avian Biology e01474 doi: 10.1111/jav.01474
  • Fallon, J.A., E.P. Smith. N. Shoch, J.D. Paruk, E.A. Adams, D.C. Evers, P.G.R. Jodice, C. Perkins, S. Shulte, W.A. Hopkins. 2017. Hematological indices of physiological injury to lightly oiled birds from the Deepwater Horizon oil spill. Environmental Toxicology and Chemistry.
  • Lamb, J.S.*, Y.G. Satge^, P.G.R. Jodice. 2017. Diet composition and provisioning rates of nestlings determine reproductive success in a subtropical seabird. Marine Ecology Progress Series 581:149-164.
  • Lamb, J.S., Y.G. Satge, P.G.R. Jodice. 2017. Influence of density-dependent competition on foraging and migratory behavior of a subtropical colonial seabird. Ecology and Evolution 2017;00:1–13. Publisher Website | 
  • Haney, J.C., P.G.R. Jodice, W.A. Montevecchi, D.C. Evers. 2017. Challenges to oil spill assessment for seabirds in the deep ocean. Archives of Environmental Contamination and Toxicology. doi:10.1007/s00244-016-0355-8
  • Zinsser, L., F. Sanders, and P. Jodice. Daily survival rate and habitat characteristics of nests of Wilson’s Plover. Southeastern Naturalist 16:149-156.
  • Ramos, R., N. Carlile, J. Madeiros, I. Ramirez, V. Paiva, H. Dinis, F. Zino, M. Biscoito, G. Leal, L. Bugoni, P.G.R. Jodice, P.G. Ryan, J. Gonzalez-Solis. 2017. The time for oceanic seabirds: tracking year-round distribution of gadfly petrels across the Atlantic Ocean. Diversity and Distributions.
  • Poli, C.L., A-L. Harrison, A. Vallarino, P.D. Gerard, Jodice, P.G.R. 2017. Dynamic oceanography predicts fine scale foraging behavior of Masked Boobies in the Gulf of Mexico. PLoS ONE 12(6): e0178318.
  • Lamb, J.S., Y.G. Satge, C.V. Fiorello, P.G.R. Jodice. 2016. Behavioral and reproductive effects of bird-borne data logger attachment on Brown Pelicans on three temporal scales. Journal of Ornithology. doi:10.1007/s10336-016-1418-3
  • Lamb, J.S.*, K.M. O’Reilly, P.G.R. Jodice. Physical condition and stress levels during early development reflect feeding rates and predict pre- and post-fledging survival in a nearshore seabird. Conservation Physiology 4 (1): cow060. doi: 10.1093/conphys/cow060
  • Collins, S.A., F.J. Sanders, and P.G.R. Jodice. Assessing conservation tools for an at-risk shorebird: Feasibility of headstarting for American Oystercatchers. Bird Conservation International 26:451-465.
  • Jodice, P.G.R., R.A. Ronconi, E. Rupp, G.E. Wallace, Y. Satge. 2015. First satellite tracks of the endangered Black-capped Petrel. Endangered Species Research 29:23-33. doi: 10.3354/esr00697 Abstract |  Download  | 
  • Lucas, J.S., S.C. Loeb, and P.G.R. Jodice. 2015 Roost selection by Rafinesque’s big-eared bats (Corynorhinus rafinesquii): a multi-scale approach. Acta Chiroptologica 17:131-141.
    Abstract | 
  • Jodice, P.G.R., J. Thibault, S.A.Collins, M. Spinks, F.J. Sanders. 2014. Reproductive ecology of American Oystercatchers nesting on shell rakes. Condor 116:588-598. Abstract | 
  • Brooks, G.L.*, F.J. Sanders, P.D. Gerard, and P.G.R. Jodice. Daily survival rates for nests of Black Skimmers from a core breeding area of the Southeastern USA. Wilson Journal of Ornithology 126:443-450. Abstract |  Download  | 
  • Ferguson, L.M.*, T.M. Norton, A. Dipl, C. Cray, M. Olivia, P.G.R. Jodice. 2014. Health assessments of brown pelican nestlings from colonies in South Carolina and Georgia. Journal of Zoo and Wildlife Medicine 45:835-845. Abstract | 
  • Carlson-Bremer, D., T.M. Norton, F.J. Sanders, B. Winn, M. Spinks, B.A. Glatt, L. Mazzaro, P.G.R. Jodice, T. Chen, E.S. Dierenfeld. 2014. Circulating fat-soluble vitamin concentrations and nutrient composition of aquatic prey eaten by American Oystercatchers in the southeastern United States. Submitted Journal of Avian Medicine and Surgery 28:216-224 Abstract | 
  • Hackney, A., R. Baldwin, P.G.R. Jodice. 2013. Mapping risk for nest predation on a barrier island. Journal of Coastal Conservation. DOI 10.1007/s11852-013-0260-5. Journal of Coastal Conservation.
  • Giles, M.M., P.G.R. Jodice, R.F. Baldwin, J. Stanton, M. Epstein. 2013. Spring migratory pathways and migration chronology of Canada Geese wintering at the Santee NWR. Canadian Field Naturalist 127: 17-25.
  • Brooks, G.L., F.J. Sanders, P.D. Gerard, P.G.R. Jodice. 2013. Predation and flooding reduce nesting success of Least Terns at natural nest sites in South Carolina. Waterbirds 36:1-10. Download  | 
  • Vander Pol, S.S., D.W. Anderson, P.G.R. Jodice, and J.E. Stuckey. 2012. East versus west: organic contaminant differences in Brown Pelican eggs from South Carolina, USA and the Gulf of California, Mexico. Science and the Total Environment 438: 527-532 Download  | 
  • Jodice, P.G.R., L.C. Wickliffe*, E.B. Sachs*. 2011. Seabird use of discards from a nearshore shrimp fishery in the South Atlantic Bight, USA. Marine Biology 158:2289-2298. Abstract | 
  • Jodice, P.G.R., and R.M. Suryan. 2010. The transboundary nature of seabird ecology. Pages 139-165In Landscape Scale Conservation Planning (R. Baldwin and S. Trembulak, eds.). Springer Verlag. Abstract | 
  • Wickliffe, L.C.*, P.G.R. Jodice. 2010. Abundance of nearshore seabirds at shrimp trawlers in South Carolina. Marine Ornithology 38:31-39. Publisher Website | 
  • Hand, C.*, F. Sanders, P.G.R. Jodice. 2010. Foraging proficiency during the nonbreeding season in a specialized forager: Are juvenile American Oystercatchers (Haematopus palliatus) 'bumble-beaks'* compared to adults? Condor 112:670-675.
  • Meehan, K.*, P.G.R. Jodice. 2010. Landscape scale correlates of fox squirrel (Sciurus niger) presence on golf courses in coastal South Carolina. Southeastern Naturalist 9:573-586.
  • Thibault, J.*, F. Sanders, P.G.R. Jodice. 2010. Parental attendance and brood success in American Oystercatchers. Waterbirds 33:511-517.
  • Sachs, E. B., and P. G. R Jodice. 2009. Behavior of parent and nestling brown pelicans during early chick rearing. Waterbirds 32:276-281
  • Eggert, L.M.F. *, P.G.R. Jodice, K.M. O’Reilly. 2009. Stress response of Brown Pelican nestlings to ectoparasite infestation. General and Comparative Endocrinology. 166: 33-38.
  • Peden-Adams, M. M., J. E. Stuckey, K. Gaworecki, J. Berger-Ritchie, K. Bryant?z P. G. R. Jodice, T. R. Scott, S. Boone, W. D. McGuinn, J. C. DeWitt, D. E. Keil. 2009. Developmental Toxicity of Perfluorooctane Sulfonate (PFOS) In White Leghorn Chickens Following In Ovo Exposure. Reproductive Toxicology 27:307-318
  • Friebel, B. A., P. G. R. Jodice. 2009. Home range and habitat use of wild hogs in the Congaree National Park, South Carolina. Human-Wildlife Conflicts 3:49-63
  • Jodice, P. G. R., D. D. Roby, K. R. Turco, R. M. Suryan, D. B. Irons, J. F. Piatt, M. T. Shultz, D. G. Roseneau, A. B. Kettle. 2008. Growth rates of Black- legged Kittiwake chicks in relation to delivery rate, size and energy density of meals. Marine Ornithology 36:107-114 Publisher Website | 
  • Eggert, L. M. F., P. G. R. Jodice. 2008. Growth of Brown Pelican nestlings exposed to sublethal levels of soft tick infestation. Condor 1104:134-142.
  • Jodice, P. G. R., T. M. Murphy, F. J. Sanders, L. Ferguson. 2007. Longterm Trends in Nest Counts of Colonial Nesting Seabirds in South Carolina, USA. Waterbirds 30:40-51
  • Jodice, P. G. R., D. D. Roby, K. R. Turco, R. M. Suryan, D. B. Irons, J. F. Piatt, M. T. Shultz, D. G. Roseneau, A. B. Kettle, and J. A. Anthony. 2006. Assessing the nutritional stress hypothesis: the relative influence of diet quantity and quality on seabird productivity. Marine Ecology Progress Series 325:267-279.
  • Suryan, R. M., D. B. Irons, E. D. Brown, P. G. R. Jodice, and D. D. Roby. 2006. Site-specific effects on productivity of an upper trophic-level marine predator: Bottom-up, top-down, and mismatch effects on reproduction in a colonial seabird. Progress in Oceanography 68:303-328.
  • Jodice, P. G. R., D. M. Epeprson, and G. H. Visser. 2006. Daily energy expenditure in free-ranging gopher tortoises (Gopherus polyphmeus). Copeia 2006:129-136.
  • Jodice, P. G. R., D. D. Roby, R. Suryan, D. B. Irons, K. R. Turco, E. D. Brown, J. F. Thedinga, and G. H. Visser. 2006. Increased energy expenditure by a seabird in response to higher food abundance. Marine Ecology Progress Series. 306:283-293.
  • Ferguson, L. M., P. G. R. Jodice, W. Post, F. I. Sanders. 2005. Reddish Egret extends its breeding range on the North American Atlantic Coast. Waterbirds 28:525-526.
  • Jodice, P. G. R., D. D. Roby, R. Suryan, D. B. Irons, G. H. Visser, M. Kaufmann, and K. R. Turco. 2003. Variation in energy expenditure among black-legged kittiwakes: effects of activity-specific metabolic rates and activity budgets. Physiological and Biochemical Zoology 76:375-388.
  • Jodice, P. G. R., D. D. Roby, D. Lyons, S. K. Nelson, S. Wright, M. Antollos, C. Anderson, S. Anderson, A. Gall, D. Rizzolo. 2003. Book review: Biology of Marine Birds. Auk 121:240-245.
  • Jodice, P. G. R., D. D. Roby, V. A. Gill, S. A. Hatch, R. B. Lanctot, and G. H. Visser. 2002. Does food availability constrain energy expenditure of black-legged kittiwakes raising young? A supplemental feeding experiment. Canadian Journal of Zoology 80:214-222.
  • Suryan, R. M., D. B. Irons, M. Kaufman, P. G. R. Jodice, D. D. Roby, J. Benson, and E. D. Brown. 2002. Short-term fluctuations in forage fish availability and the effect on prey selection and brood-rearing in the black-legged kittiwake. Marine Ecology Progress Series 236:273-287.
  • Jodice, P. G. R., S. L. Garman, and M. W. Collopy. 2001. Using resampling to assess reliability of audio-visual survey strategies for Marbled Murrelets at inland forest sites. Waterbirds 24:331-344.
  • Jodice, P. G. R., R. B. Lanctot, V. A. Gill, D. D. Roby, and S. A. Hatch. 2000. Sexing black-legged kittiwakes by DNA, behavior, and morphology. Waterbirds 23:407-417.
  • Jodice, P. G. R., and M. W. Collopy. 2000. Activity patterns of Marbled Murrelets in douglas-fir old-growth forests of the Oregon Coast Range. Condor 102:275-285.
  • Jodice, P. G. R., and M. W. Collopy. 1999. Diving and foraging patterns of Marbled Murrelets: Testing predictions from optimal breathing models. Canadian Journal of Zoology 77:1409-1418.
  • Jodice, P. G. R. 1997. Book review: Ecology and conservation of the Marbled Murrelet. Colonial Waterbirds 20: 370-372.
  • Jodice, P.G.R., S.R. Humphrey. 1993. Activity and diet of an urban population of Big Cypress fox squirrels: A reply. Journal of Wildlife Management 57:930-933.
  • Jodice, P.G.R. 1993. Movements of translocated Big Cypress fox squirrels. Florida Scientist 56:1-6.
  • Jodice, P.G.R., S.R. Humphrey. 1992. Activity and diet of an urban population of Big Cypress fox squirrels. Journal of Wildlife Management 56:685-692.
  • Humphrey, S.R., P.G.R. Jodice. 1992. Big Cypress fox squirrel. Pp. 224-233 In S.R. Humphrey, ed., Rare and endangered biota of Florida. Vol. 1. Mammals. University Press of Florida, Gainesville, FL.
  • Jodice, P.G.R. 1992. Distribution of wintering loons in the northeastern Gulf of Mexico. Pp. 172-193 In L. Morse, S. Stockwell, and M. Pokras, eds. The loon and its ecosystem: Status, management, and environmental concerns. U.S. Fish and Wildlife Service.

Current Staff

Federal Staff: 2

Masters Students: 4

Phd Students: 1

Post Docs: 1

University Staff: 2

5 Year Summary

Students graduated: 13

Scientific Publications: 25

Presentations: 82


Contact Us

South Carolina Cooperative Fish and Wildlife Research Unit Clemson, SC 29634 Phone: (864) 656 - 0168 Fax: (864) 656 - 3304

Unit Leader

Patrick Jodice
Patrick Jodice at Exumas Land and Sea Park, Bahamas

My research is focused on three primary areas: (1) breeding and foraging ecology of seabirds and shorebirds, (2) wildlife conservation physiology, and (3) wildlife ecology in urban and natural systems. I have conducted research and monitoring programs in a variety of marine and terrestrial ecosystems and have studied both avian and mammalian populations. All of my research has been driven by management and conservation issues, although the results are often of both an applied and basic nature.

My current and prior work has emphasized wildlife conservation with emphases in marine ornithology, avian ecology, reproductive energetics, foraging ecology, survey design and methodology, and urban ecology.


  1. Clemson University

South Carolina Cooperative Fish and Wildlife Research Unit Cooperators

  1. Clemson University
  2. South Carolina Department of Natural Resources
  3. U.S. Fish and Wildlife Service
  4. U.S. Geological Survey
  5. Wildlife Management Institute